This is our editor's choice of the best images added to our collection in September and October 2012. There was some great new material: rare wild Amur tigers from Vladimir Medvedev; dramatic sharks from new contributor Cheryl-Samantha Owen; great new dolphins from Brandon Cole, Tui de Roy's comprehensive Galapagos coverage; Sandra Bartocha's creative flower and landscape work; Solvin Zankl's innovative insect portraits using digital focus stacking; Anup Shah's new Kenyan material including a lion/waterbuck confrontation; British wildlife from Andy Rouse, including brown hares in action; unusual reptile and amphibian portraits from Michael Kern;
George Sanker / Black bear (Ursus americanus) young cub portrait, Yellowstone National Park, Wyoming, USA
Matthew Maran / Canada goose (Branta canadensis) gosling portrait, Hornsey, London, UK May
Cheryl-Samantha Owen / Mountain gorilla (Gorilla beringei) resting on back yawning, Bwindi Impenetrable Forest, Uganda, East Africa
Anup Shah / African lion (Panthera leo) male head and shoulders profile portrait, Masai Mara National Reserve, Kenya, September
Andy Rouse / European otter (Lutra lutra) running, UK, taken in controlled conditions July
Steven Kazlowski / Moose (Alces alces) newborn calves playing in spring vegetation. Tony Knowles Coastal Trail, Anchorage, south-central Alaska.
Ben Hall / Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire,UK, January. 'Animal Portraits' category, British Wildlife Photographer Awards (BWPA) competition 2012.
Andy Rouse / 2020VISION / Gannet (Morus bassanus) portrait showing blue eyes. Bempton Cliffs RSPB reserve, UK, Yorkshire, July.
Tui De Roy / Wolf-Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos. Did you know? Galapagos tortoises have a mutualistic relati...
Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia
Tui De Roy / Hood island giant tortoise (Chelonoidis nigra hoodensis) used for breeding program since 1960s, Tortoise Breeding Centre, Puerto Ayora, Santa Cruz Island, Galapagos Islands, Ecuador, Critically endang...
Michael D. Kern / Waxy monkey tree frog (Phyllomedusa sauvagii) captive, from Central and South America
Michael D. Kern / Madagascar Ground Gecko / Big headed gecko(Paroedura Pictus) captive, from Madagascar
Vladimir Medvedev / Camera trap image of wild Siberian tiger (Panthera tigris altaica) walking through woodland, Lazovsky Zapovednik Nature Reserve, Primorsky Krai, Far East Russia, November 1994
Michael D. Kern / Panther chameleon (Furcifer pardalis) coloured blue, walking along branch, captive, from Madagascar
Solvin Zankl / Jet Ant / Shining Jet Black Ant (Lasius fuliginosus), worker. Specimen photographed using digital focus stacking
Solvin Zankl / Close up of ant (Harpegnathos venator) head from Asia. Specimen photographed using digital focus stacking
Sandra Bartocha / Forest view of trunks of Scots Pine (Pinus sylvestris), Downy European white birch (Betula pubescens) and European Beech (Fagus sylvatica) trees, Muritz National Park, Germany, November
Sandra Bartocha / Field of ripe barley with green barley growing in vehicle tracks (Hordeum vulgare) Germany, July
Sandra Bartocha / European Beech (Fagus sylvatica) trees, with sunlight shining between branches, Germany, September
Sandra Bartocha / Waves of Baltic Sea washing over and polishing coloured pebbles, Markgrafenheide, Mecklenburg-Vorpommern, Germany, January
Sandra Bartocha / Lily of the Valley (Convallaria majalis) Stadtkreis Neubrandenburg, Mecklenburg-Vorpommern, Germany, May
Sandra Bartocha / Sporangia of moss, Dueodde, Hovedstaden, Denmark, June
Sandra Bartocha / Spring Snowflake (Leucojum vernum) in flower, Hungary, March
Sandra Bartocha / Common Poppy (Papaver rhoeas) flowers in cereal field, April
Cheryl-Samantha Owen / Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Jordi Chias / Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011, Commended in Wildlife Photographer of the Yea...
Vladimir Medvedev / Camera trap image of wild female Siberian tiger (Panthera tigris altaica) scent marking the trunk of a Cedar (Cedrus) tree, Lazovsky Zapovednik Nature Reserve, Primorsky Krai, Far East Russia, April...
Vladimir Medvedev / Camera trap image of wild male Siberian tiger (Panthera tigris altaica) scent marking the trunk of a cedar (Cedrus) tree, Lazovsky Zapovednik Nature Reserve, Primorsky Krai, Far East Russia, December...
Steven Kazlowski / Moose (Alces alces) cow with a newborn calves grazing on spring grass. Tony Knowles Coastal Trail, Anchorage, south-central Alaska, May.
Steven Kazlowski / Moose (Alces alces) cow with a newborn calf. Tony Knowles Coastal Trail, Anchorage, south-central Alaska, May.
Steven Kazlowski / Moose (Alces alces) cow suckling her newborn calves. Tony Knowles Coastal Trail, Anchorage, south-central Alaska, May.
Mark Hamblin / 2020VISION / Pine marten (Martes martes) two playing on fallen pine log in woodland, Beinn Eighe National Nature Reserve, Wester Ross, Scotland, UK
Terry Whittaker / 2020VISION / European Badger (Meles meles) on the North Downs above Folkestone. Kent, UK, June. Camera trap photo.
Loic Poidevin / Yellow warbler (Dendroica petechia) at nest, Pointe Pelee, Ontario, Canada, May