This is our editor's choice of the best images added to our collection in September and October 2012. There was some great new material: rare wild Amur tigers from Vladimir Medvedev; dramatic sharks from new contributor Cheryl-Samantha Owen; great new dolphins from Brandon Cole, Tui de Roy's comprehensive Galapagos coverage; Sandra Bartocha's creative flower and landscape work; Solvin Zankl's innovative insect portraits using digital focus stacking; Anup Shah's new Kenyan material including a lion/waterbuck confrontation; British wildlife from Andy Rouse, including brown hares in action; unusual reptile and amphibian portraits from Michael Kern;
George Sanker / Black bear (Ursus americanus) young cub portrait, Yellowstone National Park, Wyoming, USA
Matthew Maran / Canada goose (Branta canadensis) gosling portrait, Hornsey, London, UK May
Cheryl-Samantha Owen / Mountain gorilla (Gorilla beringei) resting on back yawning, Bwindi Impenetrable Forest, Uganda, East Africa
Anup Shah / African lion (Panthera leo) male head and shoulders profile portrait, Masai Mara National Reserve, Kenya, September
Andy Rouse / European otter (Lutra lutra) running, UK, taken in controlled conditions July
Steven Kazlowski / Moose (Alces alces) newborn calves playing in spring vegetation. Tony Knowles Coastal Trail, Anchorage, south-central Alaska.
Ben Hall / Whooper swan (Cygnus cygnus) in snowfall. Martin Mere Wetlands Trust, Lancashire,UK, January. 'Animal Portraits' category, British Wildlife Photographer Awards (BWPA) competition 2012.
Andy Rouse / 2020VISION / Gannet (Morus bassanus) portrait showing blue eyes. Bempton Cliffs RSPB reserve, UK, Yorkshire, July.
Tui De Roy / Wolf-Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos. Did you know? Galapagos tortoises have a mutualistic relati...
Andy Rouse / Agile wallabies (Macropus agilis) sparring and fighting, Bamarru Plains, Northern Territories, Australia