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Japanese macaques are famous for their love of hot spas in winter, but photographer Yukihiro Fukuda is the first to document their journey throughout the rest of the year.

Fukuda spent twelve months in the field photographing the macaques in Japan's Jigokudani Valley, where snow covers the ground for one third of the year. His images reveal what happens to the world's most northerly primates after the snow melts - from spring among the cherry blossoms to mating season in the autumn forest.

Fukuda trained as a vet before turning his hand to nature photography and has been documenting macaques in Japan for over eight years. His images have been highly commended in the Wildlife Photographer of the Year competition.

Japanese macaque / Snow monkey {Macaca fuscata} female and young huddle together on rock in autumn woodland, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204476
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female and young huddle together on rock in autumn woodland, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} male displays in tree hoping to attract female for mating, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204522
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male displays in tree hoping to attract female for mating, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} monkey in autumn woodland, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204474
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkey in autumn woodland, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} bathing in river in autumn, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204528
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} bathing in river in autumn, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} male grooming female in courtship in autumn, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204472
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male grooming female in courtship in autumn, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} female with young in autumn woodland, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204481
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with young in autumn woodland, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young monkey amongst autumn foliage, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204486
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey amongst autumn foliage, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} male yawning - threatening display, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204489
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male yawning - threatening display, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on pine nuts in autumn, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204490
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on pine nuts in autumn, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on seeds of the Bittersweet plant, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204526
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on seeds of the Bittersweet plant, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} five-month-old young searching for food amongst fallen leaves in autumn woodland, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204480
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} five-month-old young searching for food amongst fallen leaves in autumn woodland, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on seeds of the Bittersweet plant, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204525
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on seeds of the Bittersweet plant, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild cherry blossom in spring, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204563
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild cherry blossom in spring, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flowers in spring, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204458
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flowers in spring, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young males play fighting, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204461
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young males play fighting, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} feeding on Azalea flowers in spring, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204564
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on Azalea flowers in spring, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} four-day-old newborn baby reaches up for mother's nipple, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204502
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} four-day-old newborn baby reaches up for mother's nipple, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} four-day-old newborn monkey, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204510
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} four-day-old newborn monkey, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} mother grooming four-day-old newborn baby, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204503
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} mother grooming four-day-old newborn baby, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} at one-week-old a young baby starts to show an interest in its environment, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204514
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} at one-week-old a young baby starts to show an interest in its environment, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} two-month-old baby feeding on flower in summer, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204434
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two-month-old baby feeding on flower in summer, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} monkeys grooming beside river on hot day in summer, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204435
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys grooming beside river on hot day in summer, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} monkey jumping across a river, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204433
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkey jumping across a river, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} monkeys searching for food in river, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204465
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys searching for food in river, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} groups walking along snow trail after heavy snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204549
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} groups walking along snow trail after heavy snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} groups walking along snow trail in deep snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204546
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} groups walking along snow trail in deep snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} snow crystals on monkey's forehead, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204414
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} snow crystals on monkey's forehead, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} male in snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204548
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male in snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204411
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204382
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} two monkeys grooming in hot springs, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204407
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two monkeys grooming in hot springs, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204383
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} two one-year-old females embrace each other, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204508
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two one-year-old females embrace each other, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey bathing in hot springs, scratching its head, water at 40 degrees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204441
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey bathing in hot springs, scratching its head, water at 40 degrees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} adult female bathing in hot springs tapping the water surface with hand in play, water at 40 degrees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204388
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} adult female bathing in hot springs tapping the water surface with hand in play, water at 40 degrees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey walking through thick snow on hind legs to keep hands warmer, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204402
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey walking through thick snow on hind legs to keep hands warmer, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} female with young covered in snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204494
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with young covered in snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} baby on mother's back calling out in snow storm, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204544
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} baby on mother's back calling out in snow storm, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} male walking though snow carrying baby on back, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204406
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male walking though snow carrying baby on back, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} mother and young monkey huddle together for warmth, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204450
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} mother and young monkey huddle together for warmth, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkeys playing in the snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204417
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkeys playing in the snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} two young monkey play in deep snow, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204498
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two young monkey play in deep snow, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young monkey plays with snow ball, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204532
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey plays with snow ball, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young male mounting another monkey, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204542
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young male mounting another monkey, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204447
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204422
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204427
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan
Japanese macaque / Snow monkey {Macaca fuscata} monkey feeding on the bark of a pine tree in winter, Jigokudani, Nagano, Japan - Yukihiro Fukuda
01204429
Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkey feeding on the bark of a pine tree in winter, Jigokudani, Nagano, Japan