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Heather Angel has spent many years travelling the world in search of images which illustrate the intriguing strategies employed by plants to achieve pollination. This galllery includes many images used in her book Pollination Power published by Chicago University Press with Royal Botanic Gardens Kew.

To read more about the stories behind these images, including an interview with Heather, visit our blog. And if you’d like to order a print of any of these images, there is a new Heather Angel gallery on our print site.
Marmalade hoverfly (Episyrphus balteatus) feeding on pollen on Stargazer lily (Lilium orientalis 'Stargazer') anther. Pollination occurs when pollen is transferred to next lily flower on hover... - Heather Angel
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Heather Angel / Marmalade hoverfly (Episyrphus balteatus) feeding on pollen on Stargazer lily (Lilium orientalis 'Stargazer') anther. Pollination occurs when pollen is transferred to next lily flower on hover...
Honey bees (Apis mellifera) tearing stamens to feed on pollen of Southern magnolia (Magnolia grandiflora). Surrey, England, UK. July. - Heather Angel
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Heather Angel / Honey bees (Apis mellifera) tearing stamens to feed on pollen of Southern magnolia (Magnolia grandiflora). Surrey, England, UK. July.
Dead head hoverfly (Myathropa florea) and Leaf cutter bee (Megachile centuncularis) female feeding on Californian tree poppy (Romneya coulteri) pollen. Bee transferring pollen from abdomen to stigma.... - Heather Angel
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Heather Angel / Dead head hoverfly (Myathropa florea) and Leaf cutter bee (Megachile centuncularis) female feeding on Californian tree poppy (Romneya coulteri) pollen. Bee transferring pollen from abdomen to stigma....
Common bee fly (Bombylius major) hovering whilst nectaring on Honesty (Lunaria annua), proboscis covered with pollen. Nature reserve, Surrey, England, UK. March. - Heather Angel
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Heather Angel / Common bee fly (Bombylius major) hovering whilst nectaring on Honesty (Lunaria annua), proboscis covered with pollen. Nature reserve, Surrey, England, UK. March.
Carpenter bee (Xylocopa caffra) male piercing corolla to rob nectar from Ninepin heath (Erica mammosa), failing to pollinate flower. Harold Porter Botanic Garden, Cape Fynbos, South Africa. February. - Heather Angel
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Heather Angel / Carpenter bee (Xylocopa caffra) male piercing corolla to rob nectar from Ninepin heath (Erica mammosa), failing to pollinate flower. Harold Porter Botanic Garden, Cape Fynbos, South Africa. February.
Wasp walking around a Spanish love-in-a-mist (Nigella hispanica) flower to sip nectar from ring of blue nectaries, thorax and abdomen pick up pollen from downward facing anthers that swipe over the wa... - Heather Angel
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Heather Angel / Wasp walking around a Spanish love-in-a-mist (Nigella hispanica) flower to sip nectar from ring of blue nectaries, thorax and abdomen pick up pollen from downward facing anthers that swipe over the wa...
Wasp (Vespula germanica) nectaring on Pineapple lily (Eucomis comosa). Pollen is deposited on underside of wasp's hairy body and transferred to stigma of next flower visited. Lily cultivated in ga... - Heather Angel
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Heather Angel / Wasp (Vespula germanica) nectaring on Pineapple lily (Eucomis comosa). Pollen is deposited on underside of wasp's hairy body and transferred to stigma of next flower visited. Lily cultivated in ga...
Chinese peacock butterfly (Papilio bianor) drinking nectar from a Hibiscus flower. To maintain position it flaps the wings which pick up pollen on their undersides from the stamens placed on the end o... - Heather Angel
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Heather Angel / Chinese peacock butterfly (Papilio bianor) drinking nectar from a Hibiscus flower. To maintain position it flaps the wings which pick up pollen on their undersides from the stamens placed on the end o...
Marmalade hoverfly (Episyrphus balteatus) feeding on pollen on Viper's bugloss (Echium vulgare) anther. Dolomites, Italy. June. - Heather Angel
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Heather Angel / Marmalade hoverfly (Episyrphus balteatus) feeding on pollen on Viper's bugloss (Echium vulgare) anther. Dolomites, Italy. June.
Carpenter bee (Xylocopa fimbriata) nectaring on Quickstick / Mother of cocoa tree (Gliricidia sepium). Chiapas, Mexico. - Heather Angel
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Heather Angel / Carpenter bee (Xylocopa fimbriata) nectaring on Quickstick / Mother of cocoa tree (Gliricidia sepium). Chiapas, Mexico.
Marmalade hoverfly (Episyrphus balteatus) feeding on Caucasian lily (Lilium monadelphum) pollen. Pollen transferred to other flowers via legs. Russian Caucasus, Georgia. June. - Heather Angel
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Heather Angel / Marmalade hoverfly (Episyrphus balteatus) feeding on Caucasian lily (Lilium monadelphum) pollen. Pollen transferred to other flowers via legs. Russian Caucasus, Georgia. June.
Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July. - Heather Angel
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Heather Angel / Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July.
Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April. - Heather Angel
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Heather Angel / Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April.
Butterflies and day flying moths feed from Loofah flower (Luffa cylindrica) and pick up pollen on their legs, underside of thorax and antennae, South Sichuan Bamboo Sea, China - Heather Angel
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Heather Angel / Butterflies and day flying moths feed from Loofah flower (Luffa cylindrica) and pick up pollen on their legs, underside of thorax and antennae, South Sichuan Bamboo Sea, China
Honeybee (Apis mellifera) with full pollen baskets and pollen on head, thorax and abdomen. Bee withdrawing from Snowdrop (Galanthus nivalis) flower after nectaring. In garden, Surrey, England, UK. Feb... - Heather Angel
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Heather Angel / Honeybee (Apis mellifera) with full pollen baskets and pollen on head, thorax and abdomen. Bee withdrawing from Snowdrop (Galanthus nivalis) flower after nectaring. In garden, Surrey, England, UK. Feb...
Wool carder bee (Anthidium manicatum) female entering Rusty foxglove (Digitalis ferruginea) flower. Pollen transferring to head from stamens on upper lip of flower. Cultivated in garden, Surrey Englan... - Heather Angel
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Heather Angel / Wool carder bee (Anthidium manicatum) female entering Rusty foxglove (Digitalis ferruginea) flower. Pollen transferring to head from stamens on upper lip of flower. Cultivated in garden, Surrey Englan...
Honey bee (Apis mellifera) forages on pollen in Apple (Malus domestica) flower, collecting pollen in pollen basket. In garden, Surrey, England, UK. May. - Heather Angel
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Heather Angel / Honey bee (Apis mellifera) forages on pollen in Apple (Malus domestica) flower, collecting pollen in pollen basket. In garden, Surrey, England, UK. May.
Bumblebee (Bombus sp) nectaring on Indian horse chestnut flower (Aesculus indica). Yellow spots on banner petals turn pink as nectar resource declines. Orange pollen is pick ed up on wings and thorax... - Heather Angel
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Heather Angel / Bumblebee (Bombus sp) nectaring on Indian horse chestnut flower (Aesculus indica). Yellow spots on banner petals turn pink as nectar resource declines. Orange pollen is pick ed up on wings and thorax...
Orange breasted sunbird (Anthobaphes violacea), male nectaring on Heather (Erica sp). Kirstenbosch Botanic Garden, Cape Town, South Africa. February - Heather Angel
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Heather Angel / Orange breasted sunbird (Anthobaphes violacea), male nectaring on Heather (Erica sp). Kirstenbosch Botanic Garden, Cape Town, South Africa. February
Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August. - Heather Angel
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Heather Angel / Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August.
Cape sugarbird (Promerops cafer) male nectaring on Pincushion protea (Leucospermum cordifolium). Pollen transferred from anthers to bird's head. Kirstenbosch National Botanical Garden, Cape Town,... - Heather Angel
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Heather Angel / Cape sugarbird (Promerops cafer) male nectaring on Pincushion protea (Leucospermum cordifolium). Pollen transferred from anthers to bird's head. Kirstenbosch National Botanical Garden, Cape Town,...
Wood pigeon (Columba palumbus) feeding on sweet petals of nectarless Feijoa (Acca sellowiana). Kew Gardens, London, England, UK. July. - Heather Angel
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Heather Angel / Wood pigeon (Columba palumbus) feeding on sweet petals of nectarless Feijoa (Acca sellowiana). Kew Gardens, London, England, UK. July.
Altamira oriole (Icterus gularis) nectaring on Quickstick / Mother of cocoa tree (Gliricidia sepium) flower. Chiapas, Mexico. - Heather Angel
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Heather Angel / Altamira oriole (Icterus gularis) nectaring on Quickstick / Mother of cocoa tree (Gliricidia sepium) flower. Chiapas, Mexico.
Red wattlebird (Anthochaera carunculata) nectaring on Bottlebrush (Callistemon sp). Bird has brush-tipped tongue. Quaalup Homestead near Fitzgerald River National Park, Western Australia. - Heather Angel
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Heather Angel / Red wattlebird (Anthochaera carunculata) nectaring on Bottlebrush (Callistemon sp). Bird has brush-tipped tongue. Quaalup Homestead near Fitzgerald River National Park, Western Australia.
Scarab beetle (Scarabaeoidea) pair mating inside Tulip (Tulipa micheliana) flower. Beetles covered in pollen. Uzbekistan, April. - Heather Angel
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Heather Angel / Scarab beetle (Scarabaeoidea) pair mating inside Tulip (Tulipa micheliana) flower. Beetles covered in pollen. Uzbekistan, April.
Day gecko (Phelsuma sp.) with extended tongue lapping up nectar from crown of thorns (Euphorbia milii) flower, Madagascar - Heather Angel
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Heather Angel / Day gecko (Phelsuma sp.) with extended tongue lapping up nectar from crown of thorns (Euphorbia milii) flower, Madagascar
Honey bee (Apis mellifera) foraging on Peony (Paeonia officinalis) pollen. Surrey, England, UK. May. - Heather Angel
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Heather Angel / Honey bee (Apis mellifera) foraging on Peony (Paeonia officinalis) pollen. Surrey, England, UK. May.
Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica. - Heather Angel
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Heather Angel / Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica.
Bumblebee scarab (Amphicoma vulpes) pair mating in Tulip (Tulipa armena) flower. Pollen picked up on beetles and transferred to other flowers. Near Karabel Pass, Turkey. May. - Heather Angel
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Heather Angel / Bumblebee scarab (Amphicoma vulpes) pair mating in Tulip (Tulipa armena) flower. Pollen picked up on beetles and transferred to other flowers. Near Karabel Pass, Turkey. May.
Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China. - Heather Angel
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Heather Angel / Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China.
Long-horned bee (Eucera sp.), two crawling into Wild gladiolus (Gladiolus atroviolaceus) flower to roost at night. Turkey. May. - Heather Angel
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Heather Angel / Long-horned bee (Eucera sp.), two crawling into Wild gladiolus (Gladiolus atroviolaceus) flower to roost at night. Turkey. May.
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2. - Heather Angel
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Heather Angel / Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2.
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California. - Heather Angel
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Heather Angel / Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California.
Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April. - Heather Angel
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Heather Angel / Texas bluebonnets (Lupinus texensis) and Indian paintbrush (Castilleja coccinea) amongst wildflowers in ditch beside byway. Rains triggered germinaton of annual seeds. Cuero, Texas, USA. April.
Red poppywort (Meconopsis punicea), raindrops on flower and stem. Medicinal plant in Tibet. Balang Shan, Sichuan Province, China. June. - Heather Angel
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Heather Angel / Red poppywort (Meconopsis punicea), raindrops on flower and stem. Medicinal plant in Tibet. Balang Shan, Sichuan Province, China. June.
Red wattlebird (Anthochaera carunculata) feeding on nectar of flame grevillea (Grevillea excelsior) Kings Park, Western Australia Botanic garden, Western Australia. - Heather Angel
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Heather Angel / Red wattlebird (Anthochaera carunculata) feeding on nectar of flame grevillea (Grevillea excelsior) Kings Park, Western Australia Botanic garden, Western Australia.
Black coral vine (Kennedia nigricans) flowers, pollinated by honeyeaters (flowers triggered by honeyeater, releasing the stamens, style and stigma), Western Australia Sequence 2 of 2 - Heather Angel
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Heather Angel / Black coral vine (Kennedia nigricans) flowers, pollinated by honeyeaters (flowers triggered by honeyeater, releasing the stamens, style and stigma), Western Australia Sequence 2 of 2
Black coral vine (Kennedia nigricans) flowers, pollinated by honeyeaters (showing untriggered flowers), Western Australia Sequence 1 of 2 - Heather Angel
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Heather Angel / Black coral vine (Kennedia nigricans) flowers, pollinated by honeyeaters (showing untriggered flowers), Western Australia Sequence 1 of 2
Jade vine (Strongylodon macrobotrys) in glasshouse at Kew Gardens, London, UK. Endemic to the Philippines. Pollinated by sunbirds. - Heather Angel
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Heather Angel / Jade vine (Strongylodon macrobotrys) in glasshouse at Kew Gardens, London, UK. Endemic to the Philippines. Pollinated by sunbirds.
Honey bee (Apis mellifera) with full pollen basket foraging on pollen in Caucasian peony (Paeonia daurica Pollen transferred on bee's body to other flowers. In garden, Surrey, England, UK. May. - Heather Angel
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Heather Angel / Honey bee (Apis mellifera) with full pollen basket foraging on pollen in Caucasian peony (Paeonia daurica Pollen transferred on bee's body to other flowers. In garden, Surrey, England, UK. May.
Angle shades moth (Phlogophora meticulosa) nectaring on Snowberry (Symphoricarpos  chenaultii) at night. In garden, Surrey, England, UK. September. - Heather Angel
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Heather Angel / Angle shades moth (Phlogophora meticulosa) nectaring on Snowberry (Symphoricarpos chenaultii) at night. In garden, Surrey, England, UK. September.
Bumblebee (Bombus sp) covered with pollen grains after visiting Squash (Cucurbita sp). In garden, Surrey, England, UK. September. - Heather Angel
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Heather Angel / Bumblebee (Bombus sp) covered with pollen grains after visiting Squash (Cucurbita sp). In garden, Surrey, England, UK. September.
Musquia (Musschia aurea) flower. Cultivated in glasshouse, UK. Pollinated by Lizard (Lacerta dugesii) and Madeira bumblebee (Bombus maderensis) in Madeira. Pentaramous stigma enables lizard to climb o... - Heather Angel
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Heather Angel / Musquia (Musschia aurea) flower. Cultivated in glasshouse, UK. Pollinated by Lizard (Lacerta dugesii) and Madeira bumblebee (Bombus maderensis) in Madeira. Pentaramous stigma enables lizard to climb o...
Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation. - Heather Angel
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Heather Angel / Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation.
Stigmas in female flowers of single Wax begonia (Begonia semperflorens group) mimic pollen laden stamens produced by male flowers but offer no reward to pollinators, which land on them and pollinate t... - Heather Angel
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Heather Angel / Stigmas in female flowers of single Wax begonia (Begonia semperflorens group) mimic pollen laden stamens produced by male flowers but offer no reward to pollinators, which land on them and pollinate t...
Pollen release from cyclamen flower via sonication, which mimics buzz pollination. The structure of the cyclamen flower prevents the pollen getting wet. - Heather Angel
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Heather Angel / Pollen release from cyclamen flower via sonication, which mimics buzz pollination. The structure of the cyclamen flower prevents the pollen getting wet.
Hazel (Corylus avellana) catkins dispersing pollen with female flower above on branch. Controlled condition.s Surrey, England, UK. - Heather Angel
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Heather Angel / Hazel (Corylus avellana) catkins dispersing pollen with female flower above on branch. Controlled condition.s Surrey, England, UK.
Fly (Diptera) feeding on Pink cinquefoil (Potentilla nitida). Flies are main pollinators in Dolomites, becoming active at lower temperatures than bees. Dolomites, Italy, July. - Heather Angel
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Heather Angel / Fly (Diptera) feeding on Pink cinquefoil (Potentilla nitida). Flies are main pollinators in Dolomites, becoming active at lower temperatures than bees. Dolomites, Italy, July.
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