Sergey Gorshkov / Plume of volcanic ash lit by a basaltic lava flow from the Tolbachik volcano complex, Kamchatka, Far East Russia, eruption lasted from December 22nd 2012 through to January 2nd 2013

Nick Upton / Common prawns (Palaemon serratus) and a Rock goby (Gobius paganellus) feeding on a dead Montagu's crab (Xantho hydrophilus) in a rockpool, Cornwall, England, UK, September.

Nick Upton / Close up view of a Marbled white butterfly (Melanargia galathea) nectaring on a Field scabious (Knautia arvensis) flower, with another trying to court with it, Wiltshire, England, UK, July.

Nick Upton / Slow motion cllip of a Marbled white butterfly (Melanargia galathea) nectaring on a Creeping thistle (Cirsium arvense) flower, with another trying to court with it, Wiltshire, England, UK, July.

Nick Upton / Flock of Jackdaws (Corvus mondedula) taking off from their roost site, Gloucestershire, England, UK, December.

Nick Upton / Close-up of a Dark green fritillary (Argynnis aglaja) nectaring on a Greater knapweed (Centaurea scabiosa) flower and flying off, Wiltshire, England, UK, July.

Nick Upton / Eurasian beaver (Castor fiber) feeding on grass amongst Meadow buttercups (Ranunculus acris) near a pond in a large enclosure, Devon, England, UK, June. Captive.

Sergey Gorshkov / Plume of volcanic ash lit by a basaltic lava flow from the Tolbachik volcano complex, Kamchatka, Far East Russia, eruption lasted from December 22nd 2012 through to January 2nd 2013

James Dunbar / Male Orb-weaving spider (Araneus) courting a female near a dead fly, Bristol, England, UK, September.

Nick Upton / Eurasian beaver (Castor fiber) at night, swimming, Devon Wildlife TrustÕs Devon Beaver Project enclosure, Devon, England, UK, March.

Nick Upton / Eurasian beaver (Castor fiber) at night, chewing bark and climbing along a felled tree, Devon Wildlife TrustÕs Devon Beaver Project enclosure, Devon, England, UK, March.

Nick Upton / Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.

Nick Upton / Male Eurasian beaver (Castor fiber) at night, feeding on a branch in a pond, Devon Wildlife Trust's Devon Beaver Project, Devon, UK, April.

Fred Olivier / Tracking shot from a boat of pancake ice forming, Antarctica.

Fred Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Two King penguins (Aptenodytes patagonicus) walking along the shore, shot tilts up from feet to head, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) building its nest, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.