Tony Wu / Female Northern pygmy squid (Idiosepius paradoxus) depositing eggs on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan.

Tony Wu / Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan.

Tony Wu / Japanese dwarf flying squirrel (Pteromys volans orii) gliding through snowy forest, Hokkaido, Japan.

Tony Wu / Crab-eye goby (Signigobius biocellatus), mimics crab with eye-spot markings on dorsal fins and back and forth movement. Loloata Island, Papua New Guinea.

Tony Wu / Harlequin mantis shrimp (Odontodactylus scyllarus), close up of compound eyes. Lembeh Strait, North Sulawesi, Indonesia.

Tony Wu / Japanese dwarf flying squirrel (Pteromys volans orii) male sitting in tree, portrait. Hokkaido, Japan. March

Tony Wu / Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan.

Tony Wu / Bowhead whale (Balaena mysticetus) aerial view, Sea of Okhotsk, Russia.

Tony Wu / Bowhead whales (Balaena mysticetus) in shallow water, Sea of Okhotsk, Russia.

Tony Wu / Bowhead whale (Balaena mysticetus) tail slapping,Sea of Okhotsk, Russia.

Tony Wu / Bowhead whale (Balaena mysticetus), group socialising in shallow water, aerial view. Part of larger group of more than 50 whales congregating, Sea of Okhotsk sub-population, Russia.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female with suckling calf. Vava'u Islands, Tonga.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) male calf playing at ocean surface after coming up to breathe. Vava'u Islands, Tonga.

Tony Wu / Sperm whale (Physeter macrocephalus), ten swimming together in gathering of more than 50 individuals. Whales with flatulence and defecating. Sri Lanka.

Tony Wu / Humphead parrotfish (Bolbometopon muricatum) shoal spawning in early morning, part of mass aggregation of thousands of fish. Caroline Islands, Palau.

Tony Wu / Fringed blenny (Chirolophis japonicus) guarding its eggs. Miyagi Prefecture, Japan.

Tony Wu / Blacktip reef shark (Carcharhinus melanopterus) beaching itself to catch sardines. Tonga South Pacific.

Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.

Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...

Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.

Tony Wu / Sardines (Clupeidae) fleeing from predators in the early morning. They were being herded and chased by Trevallies and Blacktip reef sharks in the water, and picked off by brown boobies and terns from...

Tony Wu / Plastic bottles and other rubbish, found on a beach in Yamaguchi Prefecture, Japan.

Tony Wu / Humpback whale (Megaptera novaeangliae) lying on its side while executing a series of pectoral fin slaps on ocean surface. Vava'u, Tonga, South Pacific.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female breaching. Vava'u, Tonga. Pacific Ocean.

Tony Wu / Humpback whale calf (Megaptera novaeangliae australis) female nudging her mother's mammary gland whilst trying to feed. Vava;u, Tonga, South Pacific.

Tony Wu / North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching, Icy Strait, Alaska, USA. July.

Tony Wu / Sperm whale (Physeter macrocephalus) mature female, Dominica.

Tony Wu / Blue whale (Balaenoptera musculus indica) in Indian Ocean, Mirissa, Matara, Sri Lanka.

Tony Wu / Indo-Pacific sailfish (Istiophorus platypterus) in feeding frenzy, with the remains of a previously large school of sardines (Sardina sp) in background. Isla Mujeres, Cancun, Mexico.

Tony Wu / Asian sheepshead wrasse (Semicossyphus reticulatus), male in white breeding colouration. Niigata Prefecture, Honshu, Japan. June.

Tony Wu / Humpback whale (Megaptera novaeangliae australis) female, stationary with calf heading to surface for breath of air. Vava'u, Tonga, South Pacific.

Tony Wu / Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan. Highly commended in the Animal Portraits Category of the Wildlife Photographer of the Year Awa...

Tony Wu / Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image

Tony Wu / Humpback whales (Megaptera novaeangliae) male during a competetive heat run. Vava'u, Tonga, South Pacific.

Tony Wu / Bryde's whale (Balaenoptera edeni edeni) adult female feeding at surface on anchovies with her calf. Gulf of Thailand, Pacific.

Tony Wu / Humpback whale (Megaptera novaeangliae) adult female at rest. The tubercles on the whale's head are clearly visible, as are the barnacles (Coronula diadema), whale lice (Cyamus boopis), and hairs on t...

Tony Wu / Polar bear (Ursus maritimus) female with a single young cub, only a few months old, northern Svalbard, Norway, June

Tony Wu / Southern right whale (Eubalaena australis) male brindle calf at surface, born with white and black pigmentation, photographed with the permission of the Department of Environmental Affairs, South Afri...

Tony Wu / Killer whale / Orca (Orcinus orca) large adult male stalking a large school of Herring (Clupea harengus) in shallow water, Norway, February

Tony Wu / Killer whale / Orca (Orcinus orca) mature male in the foreground, with other mature males visible behind, Norway, January

Tony Wu / Humpback whale (Megaptera novaeangliae) female breaching together with her male calf. The calf had a large wound on his left peduncle area, one that appeared to be the result of a large bite, Vava'u,...

Tony Wu / Humpback whale (Megaptera novaeangliae) executing a dramatic tail slap, throwing up a cascade of water with its powerful fluke, Vava'u, Tonga, South Pacific

Tony Wu / Humpback whales (Megaptera novangliaea) unusual scene with two adults tail-slapping together in rough seas, Vava'u, Tonga, South Pacific

Tony Wu / Humpback whale (Megaptera novaeangliae) male breaching in front of Hunga island in Vava'u, Tonga, South Pacific

Tony Wu / Humpback whales (Megaptera novaeangliae) bubble net feeding with mouths open at surface, example of cooperative hunting, Chatham Strait, Alaska, USA July

Tony Wu / Humpback whale (Megaptera novangliaea) executing an energetic tail slap, creating a cascade of seawater, Vava'u, Tonga, South Pacific

Tony Wu / Humpback whale (Megaptera novaeangliae) male calf with his mother. Vava'u, Tonga. South Pacific.