Simon Littlejohn / Two European otters (Lutra lutra) hunting in the River Stour, Dorset, England, UK, May.

Simon Littlejohn / Juvenile Lammergeier vulture (Gypaetus barbatus) in flight, mobbed by a raven, Grand Bargy cirque, Haute Savoie, France, October.

Simon Littlejohn / Bonelli's eagle chicks (Aquila fasciata) in nest pecking each other, aged one week, Andalusia , Spain, May.

Simon Littlejohn / Pair of Griffon vultures (Gyps fulvus) mating, Andalusia, Spain, June.

Simon Littlejohn / Honey bee (Apis) nectaring from a flowering Eucalyptus tree, Andalusia, Spain, June.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonia) feeding on a Guanaco (Lama guanacoe) carcass, Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) dragging the carcass of a young Guanaco (Lama guanicoe), Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Close-up of a female Patagonian puma (Puma concolor patagonica) feeding on a Guanaco (Lama guanicoe) carcass, filmed with a remote camera, Torres del Paine National Park, Patagonia, Chile, May.

Simon Littlejohn / Flock of Griffon vultures (Gyps fulvus) feeding, Andalusia, Spain, May.

Simon Littlejohn / Griffon vulture (Gyps fulvus) regurgitating food for its chick, chick struggles to eat, Andalusia, Spain, June.

Simon Littlejohn / Mute swans (Cygnus olor) hissing and lunging at a European otter (Lutra lutra), River Stour, Dorset, England, UK, June.

Simon Littlejohn / Family of European otters (Lutra lutra) feeding and playing amongst Whitewater crowfoot (Ranunculus aquatilis) flowers, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Close-up of a female European otter (Lutra lutra) and two juveniles feeding in a river, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Female European otter (Lutra lutra) and two large pups playing and feeding in mid river, River Stour, Dorset, England, UK, June.

Simon Littlejohn / Kingfisher (Alcedo atthis) diving into river from perch, resurfacing with small fish and flying to branch to stun it before swallowing it, River Stour, Dorset, UK.

Simon Littlejohn / Two Otters (Lutra lutra) diving and feeding in the grass and reeds at the rivers edge, River Stour, Dorset, UK, September.

Simon Littlejohn / Grey heron (Ardea cinerea) standing on rivers edge in heavy rain, swallowing a small Pike (Esox lucius),River Stour, Dorset, UK, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) standing with its pup (approximately 14 weeks old) on the edge of the river, the pup raises its front paws onto mum's neck and the female then heads into the...

Simon Littlejohn / Adult female European otter (Lutra lutra) guarding and grooming its pup on riverbank as it eats a fish, River Stour, Dorset, UK, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) sat on riverbank grooming while her pup feeds on a fish, she then walks over to pup and grooms it before then also feeding on half of the fish, River Stour, D...

Simon Littlejohn / Adult female European otter (Lutra lutra) lying down on riverbank grooming as her pup eats a fish. The female then enters the water and swims away, River Stour, Dorset, UK, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) feeding on a fish on the river bank when her young pup arrives, takes it off her and begins feeding, River Stour, Dorset, UK, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) swimming in churned up white water near a weir, feeding on fish with her young pup at her side, Stour River Dorset, UK, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) returning to the river bank to give her young pup food, she returns to the water, shortly followed by the pup, River Stour, Dorset, UK, September..

Simon Littlejohn / European otter (Lutra lutra) pup standing on small island of vegetation in the middle of the river, awaiting the return of its mother, the female appears behind the pup who then slips into the water t...

Simon Littlejohn / Adult female European otter (Lutra lutra) swimming with her young pup attempting to ride on her back before they both dive under, River Stour, Dorset, UK, September.

Simon Littlejohn / Eurasian griffon vulture (Gyps fulvus) soaring over the valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Adult Lammergeier or Bearded vulture (Gypaetus barbatus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Adult Lammergeier or Bearded vulture (Gypaetus barbatus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Adult Lammergeier or Bearded vulture (Gypaetus barbatus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Eurasian griffon vulture (Gyps fulvus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Eurasian griffon vulture (Gyps fulvus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Bearded vulture (Gypaetus barbatus) soaring over the high valleys of the Monte Perdido y Ordessa National Park, Huesca, Aragon, Spain, September.

Simon Littlejohn / Adult female European otter (Lutra lutra) standing in shallow water by river bank with her two pups (approximately 12 weeks old), one pup is restingo n her back until she dives under the water and the...

Simon Littlejohn / Adult female European otter (Lutra lutra) fishing with her two pups (approximately 12 weeks old) following her as she clambers out on to riverbank with a small fish to feed them, River Stour, Dorset,...

Simon Littlejohn / Two European otter (Lutra lutra) pups (approximately 12 weeks old) swimming beside their mother who is diving for food, River Stour, Dorset, UK, October.

Simon Littlejohn / Two European otter (Lutra lutra) pups (approximately 12 weeks old) sitting in the shallows waiting for their mother to return before they both turn and dive into the water, River Stour, Dorset, UK, Oc...

Simon Littlejohn / Red deer (Cervus elaphus) stags rutting, with one stag urinating as they fight, Bushy Park, London, UK, September.

Simon Littlejohn / Slow worm (Anguis fragilis) resting in a patch of dry heather and bracken before slithering away under a heather bush, Wareham Forest, Dorset, UK, July.

Simon Littlejohn / Stoat (Mustela erminea) running and hunting, with winter coat, Haute-Savoie, France, October.

Simon Littlejohn / Female Patagonian puma (Puma concolor Patagonia) sheltering from a blizzard in bushes,Torres del Paine National Park, Patagonia, Chile, July.

Simon Littlejohn / Lammergeier vulture (Gypaetus barbatus) soaring, Monte Perdido y Ordesa National Park, Aragon, Spain, September.

Simon Littlejohn / Three juvenile Spanish imperial eagles (Aquila adalberti) in nest, one flapping its wings, Sierra de Andujar National Park, Andalusia, Spain, July.

Simon Littlejohn / Two Patagonian puma cubs (Puma concolor patagonica) playing, Torres del Paine National Park, Chile, April.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) walking through tall grass, with cub running in front of her, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Violet dropwing dragonfly (Trithemis annulata) taking off and landing on a stick, Sierra de Andujar Natural Park, Andalusia, Spain, August.

Simon Littlejohn / Female Patagonian puma (Puma concolor patagonica) walking through tall grass with two cubs, Torres del Paine National Park, Chile, June.

Simon Littlejohn / Bonelli's eagle (Aquila fasciata) chick defecating out of nest, Andalusia, Spain, May.