Harvest mouse (Micromys minutus) cleaning its nose on Common hogweed (Heracleum sphondylium) flowerhead after release, Moulton, Northampton, UK, June.  -  Nick Upton
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Nick Upton / Harvest mouse (Micromys minutus) cleaning its nose on Common hogweed (Heracleum sphondylium) flowerhead after release, Moulton, Northampton, UK, June.
Roe deer (Capreolus capreolus) buck running past a garden shed at night, Wiltshire, UK, April. Property released.  -  Nick Upton
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Nick Upton / Roe deer (Capreolus capreolus) buck running past a garden shed at night, Wiltshire, UK, April. Property released.
Greylag goose (Anser anser) landing with its wings and head raised to chase off another from its territory in a marshland pool, Gloucestershire, UK, January.  -  Nick Upton
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Nick Upton / Greylag goose (Anser anser) landing with its wings and head raised to chase off another from its territory in a marshland pool, Gloucestershire, UK, January.
Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December.  -  Nick Upton
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Nick Upton / Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December.
Common bee fly (Bombylius major) nectaring on a Greater periwinkle flower (Vinca major), Wiltshire garden, UK, April.  -  Nick Upton
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Nick Upton / Common bee fly (Bombylius major) nectaring on a Greater periwinkle flower (Vinca major), Wiltshire garden, UK, April.
Common carder bumblebee (Bombus pascuorum) nectaring on a Yellow archangel (Lamiastrum galeobdolon) flower in a garden, Wiltshire, UK, April  -  Nick Upton
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Nick Upton / Common carder bumblebee (Bombus pascuorum) nectaring on a Yellow archangel (Lamiastrum galeobdolon) flower in a garden, Wiltshire, UK, April
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire,...  -  Nick Upton
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Nick Upton / Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire,...
White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April.  -  Nick Upton
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Nick Upton / White-lipped snail (Cepaea hortensis) crawling over an oak sleeper retaining a garden lawn with a greenhouse in the background, Wiltshire, UK, April.
Seven-spot ladybird (Coccinnella septempunctata) just emerged from hibernation, sunning on a Honeysuckle leaf with buildings in the background, Wiltshire garden, UK, April.  -  Nick Upton
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Nick Upton / Seven-spot ladybird (Coccinnella septempunctata) just emerged from hibernation, sunning on a Honeysuckle leaf with buildings in the background, Wiltshire garden, UK, April.
Honey bee (Apis mellifera) nectaring on a Crab apple (Malus sylvestnis) flower in a garden, Wiltshire, UK, April.  -  Nick Upton
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Nick Upton / Honey bee (Apis mellifera) nectaring on a Crab apple (Malus sylvestnis) flower in a garden, Wiltshire, UK, April.
Garden snail (Cornu aspersum / Helix aspersa) crawling over an oak sleeper retaining a garden lawn with buildings and a greenhouse in the background, Wiltshire, UK, April.  -  Nick Upton
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Nick Upton / Garden snail (Cornu aspersum / Helix aspersa) crawling over an oak sleeper retaining a garden lawn with buildings and a greenhouse in the background, Wiltshire, UK, April.
Red fox (Vulpes vulpes) using a trail under a fence separating a garden from surrounding woodland and meadows at night, Wiltshire, UK, April.  -  Nick Upton
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Nick Upton / Red fox (Vulpes vulpes) using a trail under a fence separating a garden from surrounding woodland and meadows at night, Wiltshire, UK, April.
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.  -  Nick Upton
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Nick Upton / Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.  -  Nick Upton
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Nick Upton / Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.  -  Nick Upton
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.  -  Nick Upton
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...  -  Nick Upton
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Nick Upton / Six-spot burnet moth (Zygaena filipendulae), Marbled white butterflies (Melanargia galathea) and a Dark green fritillary (Argynnis aglaja) nectaring on a clump of Greater knapweed flowers (Centaurea s...
Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...  -  Nick Upton
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Nick Upton / Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...
Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...  -  Nick Upton
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Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.  -  Nick Upton
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Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.  -  Nick Upton
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.  -  Nick Upton
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Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) with a GPS tracker on its back in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...  -  Nick Upton
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Nick Upton / White stork (Ciconia ciconia) parent standing beside its three developing chicks. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cotswold...
White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.  -  Nick Upton
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Nick Upton / White stork (Ciconia ciconia) nest at captive breeding colony raising chicks for UK White Stork reintroduction project at the Knepp Estate. Cotswold Wildlife Park, Oxfordshire, UK, May 2019.
Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...  -  Nick Upton
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Nick Upton / Richard Wardle feeding fish to recently hatched White stork (Ciconia ciconia) chicks in nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Esta...
Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...  -  Nick Upton
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Nick Upton / Recently hatched White stork (Ciconia ciconia) chicks begging for food in their nest. In captive breeding colony raising young birds for UK White Stork reintroduction project at the Knepp Estate. Cots...
Pair of White storks (Ciconia ciconia) mating on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for several hundreds...  -  Nick Upton
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Nick Upton / Pair of White storks (Ciconia ciconia) mating on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for several hundreds...
Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...  -  Nick Upton
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...  -  Nick Upton
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Nick Upton / Pair of White storks (Ciconia ciconia) arranging nest material on their nest, Knepp Castle Estate, Sussex, England, UK, April. This is the first recorded instance of White storks nesting in the UK for...
White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of y...  -  Nick Upton
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Nick Upton / White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of y...
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White s...  -  Nick Upton
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Nick Upton / White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White s...
White stork (Ciconia ciconia) male landing with nest material and joining his mate on their nest in an Oak tree at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of...  -  Nick Upton
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Nick Upton / White stork (Ciconia ciconia) male landing with nest material and joining his mate on their nest in an Oak tree at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of...
View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016.  -  Nick Upton
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Nick Upton / View towards Naranjo de Bulnes (Picu Urriellu) a distinctive 2519m limestone peak, Picos de Europa mountains, viewed from Arenas de Cabrales at sunset, Asturias, Spain, August 2016.
Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August.  -  Nick Upton
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Nick Upton / Mediterranean sea holly / Pyrenean eryngo (Eryngium bourgatii) flowering on montane pastureland, Covadonga, Picos de Europa, Asturias, Spain, August.
Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August.  -  Nick Upton
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Nick Upton / Pyrenean merendera / False meadow saffron (Merendera pyrenaica / Colchicum montanum) flowering on montane pastureland, Vegas de Sotres, Picos de Europa, Asturias, Spain, August.
Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August.  -  Nick Upton
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Nick Upton / Cornish heath (Erica vagans) clump flowering on montane pastureland, above the Lakes of Covadonga, at 1300m, Picos de Europa, Asturias, Spain, August.
Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.  -  Nick Upton
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Nick Upton / Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.
Ivy bee (Colletes hederae) mating ball, showing a mass of males clustered around a female recently emerged from her burrow, Wiltshire, England, UK, September.  -  Nick Upton
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Nick Upton / Ivy bee (Colletes hederae) mating ball, showing a mass of males clustered around a female recently emerged from her burrow, Wiltshire, England, UK, September.
Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.  -  Nick Upton
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Nick Upton / Two Red fox (Vulpes vulpes) cubs playing near the entrance to their den at dusk, Somerset, England, UK, May.
European eel (Anguilla anguilla) elver migrating upstream, using an eel pass on the side of a weir, Somerset Levels, England, UK, July.  -  Nick Upton
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Nick Upton / European eel (Anguilla anguilla) elver migrating upstream, using an eel pass on the side of a weir, Somerset Levels, England, UK, July.
Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.  -  Nick Upton
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Nick Upton / Male Eurasian beaver (Castor fiber) grazing at dusk, River Otter, Devon, England, UK, June.
Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.  -  Nick Upton
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Nick Upton / Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.
Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.  -  Nick Upton
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Nick Upton / Forest caterpillar hunter (Calosoma sycophanta) feeding on a Gypsy moth caterpillar (Lymantria dispar) it has killed in a Holm oak tree, Bacu Goloritze ravine, Baunei, Sardinia, Italy, May.
Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.  -  Nick Upton
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Nick Upton / Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.
Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August.  -  Nick Upton
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Nick Upton / Feral horse (Equus caballus) walking on a remote sandy beach, near Arta, Mallorca, Spain, August.
Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September.  -  Nick Upton
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Nick Upton / Female House spider (Tegenaria sp.) at the mouth of her tubular silk retreat in an old stone wall, Somerset, UK, September.
Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September.  -  Nick Upton
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Nick Upton / Running foliage spider / Spiny-legged sac spider (Liocranum rupicola), a nationally scarce species in the UK, hunting at night in an old stone wall, Wiltshire, UK, September.
Common lace weaver / Lace-webbed spider (Amaurobius similis) male in its web in an old stone wall at night, Wiltshire, UK, September.  -  Nick Upton
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Nick Upton / Common lace weaver / Lace-webbed spider (Amaurobius similis) male in its web in an old stone wall at night, Wiltshire, UK, September.
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