Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) silverback sitting on boundary wall between Volcanoes National Park and a Potato crop, looking into valley. Area to be restored to forest. Rwanda.

Fiona Rogers / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) female portrait. Jodhpur, Rajasthan, India. March.

Wim van den Heever / Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April.

Juan Carlos Munoz / Close up of horse wearing bridle, Sierra de Gredos, Avila, Castile and Leon, Spain.

Wim van den Heever / Lion (Panthera leo) male in darkness, Okavango Delta, Botswana.

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) infant male 'Gizmo' aged 2 years, portrait. Gombe National Park, Tanzania.

Cyril Ruoso / Common woolly monkey (Lagothrix lagotricha) reaching forward, Ikamaperou Sanctuary, Amazon, Peru.

Sven Zacek / Eider ducks (Somateria mollissima) males reflected in a small pond, Longyearbyen, Svalbard, Norway. June.

Pascal Pittorino / Przewalski's horse (Equus ferus przewalski) head portrait, captive, Lozere, France, December.

Richard Du Toit / Black wildebeest (Connochaetus gnou) bull silhouetted at dawn, Rietvlei Nature Reserve, Gauteng Province, South Africa.

Ole Jorgen Liodden / Polar bear (Ursus maritimus) standing on ice floe, looking at camera. Svalbard, Norway. August.

Andy Rouse / Mute swan (Cygnus olor) with wings raised defensively at sunrise, Wales, UK, April.

George Sanker / Barred owl (Strix varia) stretching wings, Corkscrew Swamp Audubon Sanctuary, Florida, USA, March.

Andrew Parkinson / Brown hare (Lepus europaeus) adult sitting in a snow covered field. Derbyshire, UK, January.

Anup Shah / Dusky leaf monkeys (Trachypithecus obscurus) in tree, one holding baby . Khao Sam Roi Yot National Park, Thailand.

David Tipling / Common frog (Rana temporaria) and spawn in pond. West Runton, North Norfolk, England, UK, March. Highly commended in the Habitat category of the BWPA Awards 2015.

Danny Green / Snow Monkey (Macaca fuscata) with snow covered paw in front of mouth, Nagano, Japan, February

Cyril Ruoso / Adult male Drill (Mandrillus leucophaeus) Pandrillus Sanctuary, Nigeria. Non-ex

Ole Jorgen Liodden / King penguins (Aptenodytes patagonicus) walking in habitat, with mountains, South Georgia, November 2008.

Martin Gabriel / Mute swans (Cygnus olor) on water, Bavaria, Germany, February.

Tui De Roy / Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered

Andy Rouse / Mountain gorilla (Gorilla beringei) female portrait, Rwanda.

Terry Whittaker / Pine marten (Martes martes) on branch, orange glow in sky behind from the lights of Inverness. Black Isle, Scotland, UK, October. Photographed by camera trap Highly commended in the Habitat category o...

Terry Whittaker / Man sat on picnic bench in park flooded by River Thames, Chertsey, Surrey, UK, February 2014.

Kim Taylor / European hornet (Vespa crabro) worker in flight, Surrey, England, August.

Markus Varesvuo / Whooper Swan (Cygnus cygnus) standing in snowfall,Hokkaido, Japan, February

Ben Cranke / Eurasian red squirrel (Sciurus vulgaris) sitting on tree branch, Finland. February.

Alex Mustard / RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt. Red S...

Mary McDonald / African lion (Panthera leo), resting on a rock, surrounded by flies, Masai Mara Game Reserve, Kenya. November.

Olga Kamenskaya / View of people standing on ice from underneath car, on the ice of Lake Baikal, Siberia, Russia, March. Property released and Model released.

Olga Kamenskaya / Man looking into ice hole dug for ice diving, Lake Baikal, Siberia, Russia, March. Model released.

Robert Thompson / European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June.

Tim Laman / Red-crowned crane (Grus japonensis) pair in a snowstorm, Hokkaido, Japan. Endangered species.

Enrique Lopez-Tapia / Mount Annapurna I (8091m) from base camp, with hikers and prayer flags silhouetted, Annapurna Sanctuary, central Nepal, November 2011.

Bruno D'Amicis / Apennine chamois (Rupicapra pyrenaica ornata) adult male in snowy mountain habitat. Endemic to the Apennine mountains. Abruzzo, Italy, November.

Edwin Giesbers / Sumatran tiger (Panthera tigris sumatrae), captive, occurs in Sumatra, Indonesia.

Erlend Haarberg / Stoat (Mustela erminea) in white winter coat. Vauldalen, Sor-Trondelag, Norway, May.

Andy Rouse / African Lion (Panthera leo) male emerging from bush, Masai Mara, Kenya

Juan Carlos Munoz / Gelada baboon (Theropithecus gelada) troop, Simien Mountains, Ethiopia, Africa

Andrew Parkinson / Gannet (Morus bassanus) chick backlit by evening light. Shetland Islands, Scotland, UK, August.

Jussi Murtosaari / Banded Demoiselle (Calopteryx splendens) male, Finland, August

David Tipling / Great Skua (Stercorarius skua) in flight low over water, Fetlar, Shetland, June

Andy Rouse,Andy Rouse / Barn Owl (Tyto alba) in hovering hunting flight. UK, February.

Andy Rouse / African Elephant (Loxodonta africana) silhoutted against setting sun. Maasai Mara, Kenya, Africa, August.

Igor Shpilenok / Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March.

Sven Zacek,Sven Zacek / Great Tit (Parus major) in flight. Southern Estonia, December.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile female 'Bokata' aged 6 years head and shoulders portrait looking at fingers, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...