Tony Heald / African buffalo (Syncerus caffer) herd showing defensive behaviour as an African lioness (Panthera leo) walks by, Mana Pools National Park, Zimbabwe. September....

Tony Heald / Lion (Panthera leo) moving away from a defensive African buffalo (Syncerus caffer) Mana Pools National Park, Zimbabwe.

Bence Mate / African buffalo (Syncerus caffer) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / African buffalo (Syncerus caffer) and Warthog (Phacochoerus africanus) at night with full moon, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / African buffalo (Syncerus caffer) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Ann & Steve Toon / Cape buffalo (Syncerus caffer) at waterhole at night with plains zebra (Equus quagga), Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Neil Aldridge / Yellow-billed oxpecker (Buphagus africanus) perched on the back of an African buffalo (Syncerus caffer) on the banks of the Chobe River, Chobe National Park, Botswana.

Neil Aldridge / African buffalo (Syncerus caffer) standing on the banks of the Chobe River, Chobe National Park, Botswana.

Neil Aldridge / Mud-covered face of bull African buffalo (Syncerus caffer) with a cateract in one eye, on the banks of the Chobe River, Chobe National Park, Botswana.

Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sands Game Reserve, South Africa. Sequence 1 of 5

Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sands Game Reserve, South Africa. Sequence 2 of 5

Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sands Game Reserve, South Africa. Sequence 3 of 5

Sergey Gorshkov / Lioness (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sands Game Reserve, South Africa. Sequence 4 of 5

Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5

Wim van den Heever / Cape Buffalo (Syncerus caffer) in darkness, Masai Mara, Kenya.

Sharon Heald / African buffalo (Syncerus caffer) low angle portrait, Chobe National Park, Botswana.

Pal Hermansen / Yellow-billed oxpecker (Buphagus africanus) on African buffalo (Syncerus caffer), Masai Mara, Kenya

Ann & Steve Toon / Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa.

Denis-Huot / Aerial view of African elephants (Loxodonta africana) and African buffalo (Syncerus caffer) at a water hole during the dry season, Tsavo East National Park, Kenya. October.

Anup Shah / Cape buffalo (Syncerus caffer) young male approaching with curiosity. Masai Mara Nationa Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Cape buffalo (Syncerus caffer) young male approaching with curiosity. Masai Mara Nationa Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Cape buffalo (Syncerus caffer) young male approaching with curiosity. Masai Mara Nationa Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya

Jabruson / African Forest Elephant (Loxodonta africana cyclotis) juvenile chasing African Forest Buffalos (Syncerus caffer nanus). Dzanga Bai, Dzanga-Ndoki National Park, Central African Republic.

Denis-Huot,Denis-Huot / Lion (Panthera leo) male feeding on a buffalo (Syncerus caffer) Masai-Mara Game Reserve, Kenya. Vulnerable species.

Denis-Huot,Denis-Huot / Lion (Panthera leo) males feeding on buffalo (Syncerus caffer) with other buffalos watching. Masai Mara, Kenya, Vulnerable species.

Denis-Huot / African buffalo (Syncerus caffer) male covered with mud after mud bathing, Nakuru National Park, Kenya

Denis-Huot / African buffalo (Syncerus caffer) herd drinking in front of Voi lodge, aerial view, Tsavo East National Park, Kenya

Denis-Huot / African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya

Anup Shah / Forest buffalo (Syncerus caffer nanus) standing and urinating in a river running through Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Lion cub (Panthera leo) trapped by herd of Cape buffalo (Syncerus caffer caffer), Masai Mara National Reserve, Kenya

Anup Shah / Lioness (Panthera leo) trying to protect cubs from charging Cape buffalo herd (Syncerus caffer caffer), Masai Mara National Reserve, Kenya

Anup Shah / Cape / African Buffalo (Syncerus caffer) approaching with curiosity, herd in the background, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Cape / African Buffalo (Syncerus caffer) approaching camera with curiosity, wide angle perspective, Masai Mara National Reserve, Kenya. September

Chris Shields / Illustration of wildlife at African waterhole: Water buffalo (Syncerus caffer),gazelle,African lion (Panthera leo),African elephant (Loxodonta sp), weaver birds (Ploceidae).

Anup Shah / Two Cape / African Buffalo (Syncerus caffer) males fighting as a test of strength, Masai Mara National Reserve, Kenya. February

Anup Shah / African lions (Panthera leo) climbing fallen tree in attempt to escape from herd of Cape Buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya. March.

Anup Shah / Cape buffalo (Syncerus caffer) head portrait of male, with Cattle egret on its back (Bubulcus ibis). Masai Mara National Reserve, Kenya. February.

Richard Du Toit / Aerial view of herd of African buffalo (Syncerus caffer) on the move, Okavango delta, Botswana

Marc Slootmaekers / A bloody lion (Panthera leo) lying on its back after feeding on a buffalo, Masai Mara, Kenya. Veolia Environnement Wildlife Photographer of the Year 2009: Animal Portraits category: Runner-up

Andy Rouse / African lion (Panthera leo) hunting Buffalo, South Luangwa, Zambia (non-ex)

Anup Shah / Cape buffalo (Syncerus caffer) at a water hole, Masai Mara National Reserve, Kenya, February

Nick Garbutt / Male African / Cape Buffalo (Syncerus caffer) Okavango Delta, Botswana.

Anup Shah / African lions {Panthera leo} bringing down an African buffalo {Syncerus caffer} Masai Mara GR, Kenya