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    • Aggression
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Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June  -  Benjamin Barthelemy,Benjamin  Barthelemy
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Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.  -  Nature Production
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Nature Production / Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.
Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS,  GDT 2011 competition  -  Ole Jorgen Liodden
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Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition
Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.  -  Markus Varesvuo
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Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.
Guadalupe fur seal (Arctocephalus townsendi) male fighting, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April  -  Claudio Contreras
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Claudio Contreras / Guadalupe fur seal (Arctocephalus townsendi) male fighting, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...  -  Ben Cranke
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Ben Cranke / African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...
Dalmatian pelican (Pelicanus crispus) group squabbling over fish, Lake Kerkini, Greece.  -  Edwin Giesbers
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Edwin Giesbers / Dalmatian pelican (Pelicanus crispus) group squabbling over fish, Lake Kerkini, Greece.
Portrait of an ant-mimicking crab spider. (Amyciaea sp) raising its legs to warn the photographer. Buxa tiger reserve, India.  -  Ripan Biswas
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Ripan Biswas / Portrait of an ant-mimicking crab spider. (Amyciaea sp) raising its legs to warn the photographer. Buxa tiger reserve, India.
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October.  -  Tony Heald
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Tony Heald / Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October.
Mute swans (Cygnus olor) hissing and lunging at a European otter (Lutra lutra), River Stour, Dorset, England, UK, June.  -  Simon Littlejohn
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Simon Littlejohn / Mute swans (Cygnus olor) hissing and lunging at a European otter (Lutra lutra), River Stour, Dorset, England, UK, June.
Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;  -  Oscar Dewhurst
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Oscar Dewhurst / Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
Mountain hare (Lepus timidus) fighting in the snow at night, Vauldalen, Norway, April.  -  Erlend Haarberg
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Erlend Haarberg / Mountain hare (Lepus timidus) fighting in the snow at night, Vauldalen, Norway, April.
Red fox cubs (Vulpes vulpes) play fighting in an allotment, London, England, UK, June.  -  Matthew Maran
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Matthew Maran / Red fox cubs (Vulpes vulpes) play fighting in an allotment, London, England, UK, June.
Wolf giant tortoise (Chelonoidis becki) two males fight briefly over mating rights. Wolf Volcano, Isabela Island, Galapagos  -  Tui De Roy
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Tui De Roy / Wolf giant tortoise (Chelonoidis becki) two males fight briefly over mating rights. Wolf Volcano, Isabela Island, Galapagos
Red Deer Stags (Cervus elaphus) fighting amongst the bracken. Bradgate Park, Leicestershire, UK, October.  -  Danny Green
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Danny Green / Red Deer Stags (Cervus elaphus) fighting amongst the bracken. Bradgate Park, Leicestershire, UK, October.
Hawfinch (Coccothraustes coccothraustes) fighting in snow, Bulgaria  -  Guy Edwardes
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Guy Edwardes / Hawfinch (Coccothraustes coccothraustes) fighting in snow, Bulgaria
Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.  -  Morley Read
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Morley Read / Slow motion clip of a Linnaeus' sipo snake (Chironius exoletus) with open mouth, part of threat display, Amazon rainforest, Orellana Province, Ecuador.
Parrot snake (Leptophis ahaetulla) threat display, Amazon rainforest, Orellana Province, Ecuador.  -  Morley Read
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Morley Read / Parrot snake (Leptophis ahaetulla) threat display, Amazon rainforest, Orellana Province, Ecuador.
Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.  -  Nick Upton
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Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...  -  Alex Mustard
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Alex Mustard / Two female Common seahorses (Hippocampus kuda) fighting over territory on the seabed. The brief fight was preceeded by a ritualised, synchronised parade. Bitung, North Sulawesi, Indonesia. Lembeh Stra...
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia.  -  Steven David Miller
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Steven David Miller / Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia.
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia.  -  Steven David Miller
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Steven David Miller / Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia.
Eastern grey kangaroo (Macropus giganteus), two males fighting, evenly matched. Grampians National Park, Victoria, Australia.  -  Steven David Miller
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Steven David Miller / Eastern grey kangaroo (Macropus giganteus), two males fighting, evenly matched. Grampians National Park, Victoria, Australia.
Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...  -  John Chan
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John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July.  -  Nick Hawkins
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Nick Hawkins / European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July.
RF - Black Grouse (Tetrao tetrix), two males fighting on lek , Scotland, UK.April (This image may be licensed either as rights managed or royalty free.)  -  SCOTLAND: The Big Picture
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SCOTLAND: The Big Picture / RF - Black Grouse (Tetrao tetrix), two males fighting on lek , Scotland, UK.April (This image may be licensed either as rights managed or royalty free.)
RF - Two Red foxes (Vulpes vulpes) on hind legs play fighting, Netherlands. (This image may be licensed either as rights managed or royalty free.)  -  Terry  Whittaker
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Terry Whittaker / RF - Two Red foxes (Vulpes vulpes) on hind legs play fighting, Netherlands. (This image may be licensed either as rights managed or royalty free.)
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.  -  Marie  Read
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Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA  -  Marie  Read
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Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.  -  Marie  Read
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Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.
Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April  -  David Tipling
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David Tipling / Common European adder (Vipera berus) two males fighting over a female. Holt, North Norfolk, England, UK. April
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April  -  David Tipling
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David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April
Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.  -  David Tipling
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David Tipling / Common European adder (Vipera berus) two males fighting over a female, Holt, North Norfolk, England, UK. April.
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May  -  David  Pattyn
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David Pattyn / Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Mustang stallion kicking out after winning fight. Rocky Mountains, USA, June.  -  Klein & Hubert
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Klein & Hubert / Mustang stallion kicking out after winning fight. Rocky Mountains, USA, June.
Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.  -  George  Sanker
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George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.  -  Tony Wu
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Tony Wu / View down the throat of a male Zoarchias major eelpout (Zoarchias major) that has opened its mouth during a confrontation with another male. Yamaguchi Prefecture, Japan. Pacific Ocean.
Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.  -  Tony Wu
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Tony Wu / Two male Zoarchias major eelpouts (Zoarchias major) engaged in a mouth-to-mouth confrontation. Yamaguchi Prefecture, Japan, Pacific Ocean.
Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...  -  Tony Wu
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Tony Wu / Zoarchias major eelpouts. (Zoarchias major) The orange individual is female, the one in the hole is male. The female has approached the male, curled around him, attempting to dislodge the male from th...
Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.  -  Tony Wu
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Tony Wu / Zoarchias major eelpout (Zoarchias major) with her mouth wide open. Yamaguchi Prefecture, Japan, Pacific Ocean.
Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.  -  Stefan Christmann
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.
Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.  -  Stefan Christmann
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
Mozambique spitting cobra (Naja mossambica) spitting venom. Kruger, South Africa. Controlled conditions.  -  Guy Edwardes
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Guy Edwardes / Mozambique spitting cobra (Naja mossambica) spitting venom. Kruger, South Africa. Controlled conditions.
Pere David's deer / Milu (Elaphurus davidianus), two stags testing their strength in Hubei Tian'ezhou Milu National Nature Reserve, Shishou, Hubei, China  -  Staffan Widstrand / Wild Wonders of China
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Staffan Widstrand / Wild Wonders of China / Pere David's deer / Milu (Elaphurus davidianus), two stags testing their strength in Hubei Tian'ezhou Milu National Nature Reserve, Shishou, Hubei, China
Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.  -  David Perpinan
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David Perpinan / Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.
Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, with another chick leaning in, chased away by adult, Barcelona, Spain, May.  -  David Perpinan
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David Perpinan / Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, with another chick leaning in, chased away by adult, Barcelona, Spain, May.
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.  -  Paul Hobson
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Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.  -  Paul Hobson
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Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
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