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RF-Gelada baboon (Theropithecus gelada) dominant male running. Debre Libanos, Rift Valley, Ethiopia. 2017. (This image may be licensed either as rights managed or royalty free)  -  Sylvain Cordier
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Sylvain Cordier / RF-Gelada baboon (Theropithecus gelada) dominant male running. Debre Libanos, Rift Valley, Ethiopia. 2017. (This image may be licensed either as rights managed or royalty free)
Gelada baboon (Theropithecus gelada) female carrying baby on back, dominant male in background. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada) female carrying baby on back, dominant male in background. Debre Libanos, Rift Valley, Ethiopia.
Mountain gorilla (Gorilla beringei beringei), dominant silverback  in forest clearing. Volcanoes National Park, Rwanda.  -  Christophe Courteau
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Christophe Courteau / Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda.
Mountain gorilla (Gorilla beringei beringei) silverback sitting on boundary wall between Volcanoes National Park and a Potato crop, looking into valley. Area to be restored to forest. Rwanda.  -  Christophe Courteau
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Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) silverback sitting on boundary wall between Volcanoes National Park and a Potato crop, looking into valley. Area to be restored to forest. Rwanda.
Meerkat (Suricata suricatta) alpha female nursing her one month old pups on the den, Kalahari Desert, Botswana.  -  Klein & Hubert
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Klein & Hubert / Meerkat (Suricata suricatta) alpha female nursing her one month old pups on the den, Kalahari Desert, Botswana.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.  -  Fiona Rogers
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Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.
Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.  -  Konrad  Wothe
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Konrad Wothe / Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.
Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.  -  Konrad  Wothe
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Konrad Wothe / Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.
Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month hanging upside down in a tree, Tangkoko National Park, Sulawesi, Indonesia.  -  Anup Shah
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Anup Shah / Celebes / Black crested macaque (Macaca nigra) 'Alpha' baby female aged about 1 month hanging upside down in a tree, Tangkoko National Park, Sulawesi, Indonesia.
Red deer (Cervus elaphus) dominant stag at rut,  Bradgate Park, Leicestershire, England, UK, November  -  Danny Green / 2020VISION
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Danny Green / 2020VISION / Red deer (Cervus elaphus) dominant stag at rut, Bradgate Park, Leicestershire, England, UK, November
Western chimpanzee (Pan troglodytes verus)   infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 4...  -  Anup Shah
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Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years acting aggressively towards alpha male 'Foaf' aged 30 years, Bossou Forest, Mont Nimba, Guinea. January 2011. Sequence 4...
African Lion pride male (Panthera leo) head portrait, low angle, Masai Mara National Reserve, Kenya. February  -  Anup Shah
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Anup Shah / African Lion pride male (Panthera leo) head portrait, low angle, Masai Mara National Reserve, Kenya. February
Alpha male Chimpanzee (Pan troglodytes) self grooming. Tropical forest, Western Uganda.  -  Suzi Eszterhas
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Suzi Eszterhas / Alpha male Chimpanzee (Pan troglodytes) self grooming. Tropical forest, Western Uganda.
Fishing vessel "Ocean Harvest" operating close to the "Beryl Alpha" production platform, 160 miles Northeast of Aberdeen, in the North Sea. March 2010.  -  Philip Stephen
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Philip Stephen / Fishing vessel "Ocean Harvest" operating close to the "Beryl Alpha" production platform, 160 miles Northeast of Aberdeen, in the North Sea. March 2010.
White handed gibbon, alpha male hanging in tree {Hylobates lar} Khao Yai NP, Thailand 'Cassius'  -  Justine Evans
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Justine Evans / White handed gibbon, alpha male hanging in tree {Hylobates lar} Khao Yai NP, Thailand 'Cassius'
Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002  -  Anup Shah
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Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002
Chimpanzees gather round alpha male 'Frodo' in tree with dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002  -  Anup Shah
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Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree with dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002
Ousted alpha male Chimpanzee 'Freud'  feeding on mangoes, {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002  -  Anup Shah
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Anup Shah / Ousted alpha male Chimpanzee 'Freud' feeding on mangoes, {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002
Alpha male 'Frodo' displays 'keep away' message to begging chimps wanting share of Bushbuck fawn prey {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002  -  Anup Shah
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Anup Shah / Alpha male 'Frodo' displays 'keep away' message to begging chimps wanting share of Bushbuck fawn prey {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002
Alpha male Chimpanzee 'Frodo', 26-years-old, feeding on leaves {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002. Chimps have learnt that certain leaves can be used to tre...  -  Anup Shah
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Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old, feeding on leaves {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002. Chimps have learnt that certain leaves can be used to tre...
Alpha male Chimpanzee 'Frodo', 26-years-old {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002  -  Anup Shah
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Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002
Drill (Mandrillus leucophaeus) alpha male calling, Pandrillus Drill Sanctuary, Cross River State, Nigeria  -  Cyril Ruoso
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Cyril Ruoso / Drill (Mandrillus leucophaeus) alpha male calling, Pandrillus Drill Sanctuary, Cross River State, Nigeria
Drill (Mandrillus leucophaeus) adult female grooming alpha male, Pandrillus Drill Sanctuary, Nigeria  -  Cyril Ruoso
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Cyril Ruoso / Drill (Mandrillus leucophaeus) adult female grooming alpha male, Pandrillus Drill Sanctuary, Nigeria
Timber Wolf (Canis lupus) alpha female resting as yearling growls at camera, Minnesota  -  Jim Brandenburg
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Jim Brandenburg / Timber Wolf (Canis lupus) alpha female resting as yearling growls at camera, Minnesota
Timber Wolf (Canis lupus) alpha pair in light snowfall, Minnesota  -  Jim Brandenburg
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Jim Brandenburg / Timber Wolf (Canis lupus) alpha pair in light snowfall, Minnesota
Arctic Wolf (Canis lupus) alpha male known as Buster, smelling Arctic Poppy (Papaver lapponicum) flowers, Ellesmere Island, Nunavut, Canada  -  Jim Brandenburg
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Jim Brandenburg / Arctic Wolf (Canis lupus) alpha male known as Buster, smelling Arctic Poppy (Papaver lapponicum) flowers, Ellesmere Island, Nunavut, Canada
Hamadryas baboon (Papio hamadryas), dominant male running. Awash National Park, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Hamadryas baboon (Papio hamadryas), dominant male running. Awash National Park, Rift Valley, Ethiopia.
Hamadryas baboon (Papio hamadryas), two, dominant male sitting and female standing. Awash National Park, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Hamadryas baboon (Papio hamadryas), two, dominant male sitting and female standing. Awash National Park, Rift Valley, Ethiopia.
Hamadryas baboon (Papio hamadryas), dominant male sitting. Awash National Park, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Hamadryas baboon (Papio hamadryas), dominant male sitting. Awash National Park, Rift Valley, Ethiopia.
Hamadryas baboon (Papio hamadryas), two, dominant male sitting with member of family. Awash National Park, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Hamadryas baboon (Papio hamadryas), two, dominant male sitting with member of family. Awash National Park, Rift Valley, Ethiopia.
Sign in Awash National Park about Hamadryas baboon (Papio hamadryas) along road. Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Sign in Awash National Park about Hamadryas baboon (Papio hamadryas) along road. Rift Valley, Ethiopia.
Gelada baboon (Theropithecus gelada), dominant male in aggressive stance. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada), dominant male in aggressive stance. Debre Libanos, Rift Valley, Ethiopia.
Gelada baboon (Theropithecus gelada), dominant male sitting on rock. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada), dominant male sitting on rock. Debre Libanos, Rift Valley, Ethiopia.
Gelada baboon (Theropithecus gelada), dominant male walking. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada), dominant male walking. Debre Libanos, Rift Valley, Ethiopia.
Gelada baboon (Theropithecus gelada) group including dominant male mating with female. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada) group including dominant male mating with female. Debre Libanos, Rift Valley, Ethiopia.
Gelada baboon (Theropithecus gelada), dominant male resting, portrait. Debre Libanos, Rift Valley, Ethiopia.  -  Sylvain Cordier
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Sylvain Cordier / Gelada baboon (Theropithecus gelada), dominant male resting, portrait. Debre Libanos, Rift Valley, Ethiopia.
Wolf (Canis lupus), Alpha male running. Kuhmo, Finland, October.  -  Danny Green
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Danny Green / Wolf (Canis lupus), Alpha male running. Kuhmo, Finland, October.
European Wolf (Canis lupus), alpha female hunting. Finland, July.  -  Danny Green
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Danny Green / European Wolf (Canis lupus), alpha female hunting. Finland, July.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Schweini' aged 23 years apologising to 'Alpha' male 'Ferdinand' aged 21 years . Gombe National Park, Tanzania. May...  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Schweini' aged 23 years apologising to 'Alpha' male 'Ferdinand' aged 21 years . Gombe National Park, Tanzania. May...
Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest watched by his brother 'Faustino' aged 25 years . Go...  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest watched by his brother 'Faustino' aged 25 years . Go...
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years responding to an incoming male . Gombe National Park, Tanzania. September 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years responding to an incoming male . Gombe National Park, Tanzania. September 2014.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years responding to an incoming male . Gombe National Park, Tanzania. September 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years responding to an incoming male . Gombe National Park, Tanzania. September 2014.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest while female 'Samwise' aged 12 begs for some . Gombe...  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest while female 'Samwise' aged 12 begs for some . Gombe...
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest watched by his brother 'Faustino' aged 25 years . Go...  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest watched by his brother 'Faustino' aged 25 years . Go...
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years resting . Gombe National Park, Tanzania. September 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years resting . Gombe National Park, Tanzania. September 2014.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years being groomed . Gombe National Park, Tanzania. May 2014.  -  Anup Shah
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years being groomed . Gombe National Park, Tanzania. May 2014.
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