David Pattyn / Mute swan (Cygnus olor) in a threathening territorial pose and attacking a rival, Lake Geneva, near Geneva, Switzerland, March
Ben Cranke / African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia
Ben Cranke / King Penguins (Aptenodytes patagonicus) approached by an Antarctic Fur Seal (Arctocephalus gazella). A few pecks from the penguins caused the seal to retreat. Southern elephant seal (Mirounga leonina)...
Sven Zacek / Ural Owl (strix uralensis) female with talons ready to attack a ringing ornithologist, Tartumaa county, Southern Estonia. May.
Dave Bevan / Slow motion clip of a Great spotted woodpecker (Dendrocopos major) returning to nest hole, attacked by a Nuthatch (Sitta europaea), Carmarthenshire, Wales, UK, April.
Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
John Chan / Western gulls (Larus occidentalis) attacking an Elegant tern (Thalasseus elegans) with a broken wing, Southern California, USA, April.
David Perpinan / Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, with another chick leaning in, chased away by adult, Barcelona, Spain, May.
Neil Aldridge / Lion cub (Panthera leo) playfully attacking its mother as she drags her puku antelope prey across burnt Busanga Plains, Kafue National Park, Zambia. Vulnerable
Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5
Andy Rouse / Bengal tiger (Panthera tigris tigris) female 'Noor T19' bringing down Sambar deer (Rusa unicolor) Ranthambhore, India. Sequence 6 of 18.
Claudio Contreras / Atlantic sailfish (Istiophorus albicans) attacking school of Sardine (Sardinella aurita) bait ball, Isla Mujeres, Caribbean Sea, Mexico, February
Wim van den Heever / Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Sequence 4 of 5
Wim van den Heever / Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015.
Sven Zacek / Ravens (Corvus corax) fighting in bog during snowfall, Tartumaa, Estonia. December.
Solvin Zankl / Black backed jackal (Canis mesomelas) attacking Cape fur seal (Arctocephalus pusillus) pup, Sperrgebiet National Park, Namibia, December. Sequence 3/6
David Pattyn / Great crested grebe (Podiceps cristatus) attacking another grebe in the mating season during a territorial dispute. The Netherlands. April.
Konrad Wothe / Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.
Konrad Wothe / Showreel of Male Chimpanzees (Pan troglodytes) attacking the alpha male of their troop, Mahale Mountains National Park, Tanzania.