Charlie Hamilton James / Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.
Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Marie Read / Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions.
Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.
John Chan / Male Anna's hummingbird (Calypte anna) cleaning its beak on a twig, Southern California, USA, January.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick begging for food, on parent's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) egg hatching on father's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Paul Hobson / Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.
Paul Hobson / Spoonbill (Platalea leucorodia) preening, close up. Hungary. May.
Kerstin Hinze / Red crossbill (Loxia curvirostra) feeding from a spruce cone, Bavarian Forest National Park, Germany, October.
Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
Sven Zacek / Ural owl (Strix uralensis) female flying with prey in beak. Flying back to nest in evening light. Tartumaa, Southern Estonia. May.
Tui De Roy / Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos.
Konrad Wothe / Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador.
Steven David Miller / Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive.
Guy Edwardes / Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) sleeping in the breeding colony, brooding egg, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of the Terr...