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Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion. - Marie  Read
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Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.
Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January. - Marie  Read
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Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Grey wolf (Canis lupus) female and two cubs, one howling. Controlled conditions - Klein & Hubert
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Klein & Hubert / Grey wolf (Canis lupus) female and two cubs, one howling. Controlled conditions
Wolf (Canis lupus) puppy, age one month, Rocky Mountains, USA. Controlled conditions. - Klein & Hubert
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Klein & Hubert / Wolf (Canis lupus) puppy, age one month, Rocky Mountains, USA. Controlled conditions.
Emperor penguin (Aptenodytes forsteri), group of three males incubating, one with begging chick. Atka Bay, Antarctica. August. - Stefan Christmann
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), group of three males incubating, one with begging chick. Atka Bay, Antarctica. August.
Male Mediterranean tree frog (Hyla meridionalis) calling, showing vocal sac inflating, Barcelona, Spain, May. Filmed at night with an infrared camera. - David Perpinan
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David Perpinan / Male Mediterranean tree frog (Hyla meridionalis) calling, showing vocal sac inflating, Barcelona, Spain, May. Filmed at night with an infrared camera.
Territorial Common raven (Corvus corax) on cliff top, tearing out vegetation, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May. - Philippe Clement
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Philippe Clement / Territorial Common raven (Corvus corax) on cliff top, tearing out vegetation, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
Male  Pied flycatcher (Ficedula hypoleuca) calling to establish territory and attract a mate, Carmarthenshire, Wales, UK, May. - Dave Bevan
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Dave Bevan / Male Pied flycatcher (Ficedula hypoleuca) calling to establish territory and attract a mate, Carmarthenshire, Wales, UK, May.
Pheasant (Phasianus colchicus) male calling,  East Frisian Islands, Wittbulten National Park.  Wadden Sea UNESCO World Heritage Site, Germany, June. - Konrad  Wothe
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Konrad Wothe / Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June.
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling  and displaying to attract female by waving foot, Agumbe, Western Ghats, India. - Yashpal Rathore
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling  and displaying to attract female by waving foot, Agumbe, Western Ghats, India. - Yashpal Rathore
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
Oystercatcher (Haematopus ostralegus) incubating eggs on nest, alarm calling as a predator flies overhead, Scotland,UK, June. - David Perpinan
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David Perpinan / Oystercatcher (Haematopus ostralegus) incubating eggs on nest, alarm calling as a predator flies overhead, Scotland,UK, June.
Emperor penguin (Aptenodytes forsteri) chick begging for food from parent, Adelie Land, Antarctica, January. - Fred  Olivier
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Fred Olivier / Emperor penguin (Aptenodytes forsteri) chick begging for food from parent, Adelie Land, Antarctica, January.
Emperor penguin (Aptenodytes forsteri) chicks calling, with another small chick arriving and falling over, Adelie Land, Antarctica, January. - Fred  Olivier
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Fred Olivier / Emperor penguin (Aptenodytes forsteri) chicks calling, with another small chick arriving and falling over, Adelie Land, Antarctica, January.
Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India. - Yashpal Rathore
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Yashpal Rathore / Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India.
Japanese / Red-crowned crane (Grus japonicus) two calling and dancing, part of bonding and courtship display, Hokkaido Japan February - Markus Varesvuo
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Markus Varesvuo / Japanese / Red-crowned crane (Grus japonicus) two calling and dancing, part of bonding and courtship display, Hokkaido Japan February
Pack of Grey wolves (Canis lupus) howling in the snow, captive. Bavarian Forest National Park, Germany, March. Captive. - Eric Baccega
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Eric Baccega / Pack of Grey wolves (Canis lupus) howling in the snow, captive. Bavarian Forest National Park, Germany, March. Captive.
RF- Lesser black-backed gulls (Larus fuscus) calling from shoreline, beaks open, England, UK, May. (This image may be licensed either as rights managed or royalty free.) - Ernie  Janes
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Ernie Janes / RF- Lesser black-backed gulls (Larus fuscus) calling from shoreline, beaks open, England, UK, May. (This image may be licensed either as rights managed or royalty free.)
Lion cub (Panthera leo) calling to its mother from the raised deck of a tourist lodge in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable species. - Neil Aldridge
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Neil Aldridge / Lion cub (Panthera leo) calling to its mother from the raised deck of a tourist lodge in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable species.
Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in  British Birds Bird Photograph of the Year competition 2016. - Mark Hamblin
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Mark Hamblin / Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016.
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica - Alex  Hyde
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Alex Hyde / Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012. - Fiona Rogers
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Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.
Glaucous gull (Larus hyperboreus) standing on edge of ice calling, Svalbard, Norway - Klein & Hubert
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Klein & Hubert / Glaucous gull (Larus hyperboreus) standing on edge of ice calling, Svalbard, Norway
Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica. - Bence  Mate
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Bence Mate / Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.
White tailed sea eagle (Halieetus albicilla) juvenile and adult calling on lake shore, Lake Csaj, Kiskunsagi National Park, Hungary. February. - Bence  Mate
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Bence Mate / White tailed sea eagle (Halieetus albicilla) juvenile and adult calling on lake shore, Lake Csaj, Kiskunsagi National Park, Hungary. February.
White tailed sea eagle (Halieetus albicilla) head portrait beak open calling,  Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January. - Bence  Mate
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Bence Mate / White tailed sea eagle (Halieetus albicilla) head portrait beak open calling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, January.
Greater roadrunner (Geococcyx californianus) with nuptial gift calling mate, Laredo Borderlands, Texas, USA. April - Claudio  Contreras
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Claudio Contreras / Greater roadrunner (Geococcyx californianus) with nuptial gift calling mate, Laredo Borderlands, Texas, USA. April
Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa. - Neil Aldridge
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Neil Aldridge / Southern yellow-billed hornbill (Tockus leucomelas) displays while calling in winter, Kruger National Park, South Africa.
Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years calling while holding her baby granddaughter aged 2 months and her own infant son aged 2 years. Gombe National Park,... - Fiona Rogers
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Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years calling while holding her baby granddaughter aged 2 months and her own infant son aged 2 years. Gombe National Park,...
Black spider monkey (Ateles chamek) calling, captive, Peru. - Mark  Bowler
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Mark Bowler / Black spider monkey (Ateles chamek) calling, captive, Peru.
Atlantic puffin (Larus tridactyla) seen from burrow, Iceland, July. - Cyril Ruoso
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Cyril Ruoso / Atlantic puffin (Larus tridactyla) seen from burrow, Iceland, July.
Japanese macaque (Macaca fuscata fuscata) rare white furred baby calling, Jigokudani Valley,  Nagano Prefecture, Japan. June. - Yukihiro  Fukuda
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Yukihiro Fukuda / Japanese macaque (Macaca fuscata fuscata) rare white furred baby calling, Jigokudani Valley, Nagano Prefecture, Japan. June.
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February. - Ernie  Janes
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Ernie Janes / Robin (Erithacus rubecula) singing, Norfolk, England, UK, February.
Bay-winged cowbird,(Agelaioides badius) chicks begging for food, La Pampa, Argentina - Gabriel Rojo
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Gabriel Rojo / Bay-winged cowbird,(Agelaioides badius) chicks begging for food, La Pampa, Argentina
Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia. - Fred  Olivier
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Fred Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.
Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia. - Fred  Olivier
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Fred Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.
Atlantic puffin (Fratercula arctica) two calling aggresively in snow, Hornya birdcliff, Finnmark, Norway. - Espen Bergersen
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Espen Bergersen / Atlantic puffin (Fratercula arctica) two calling aggresively in snow, Hornya birdcliff, Finnmark, Norway.
Common guillemot  (Uria aalge) calling, bridled guillemot morph with white ring around the eye, Hornoya bird cliff, Finnmark, Norway. March. - Espen Bergersen
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Espen Bergersen / Common guillemot (Uria aalge) calling, bridled guillemot morph with white ring around the eye, Hornoya bird cliff, Finnmark, Norway. March.
Common raven (Corvus corax) pair in courtship display, flying upsidedown and calling, Hornoya bird cliff, Norway. March. - Espen Bergersen
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Espen Bergersen / Common raven (Corvus corax) pair in courtship display, flying upsidedown and calling, Hornoya bird cliff, Norway. March.
Japanese crane (Grus japonensis) displaying, Hokkaido, Japan, February - Danny Green
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Danny Green / Japanese crane (Grus japonensis) displaying, Hokkaido, Japan, February
Red deer (Cervus elaphus) male calling during rut, Denmark. Non-ex - Cyril Ruoso
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Cyril Ruoso / Red deer (Cervus elaphus) male calling during rut, Denmark. Non-ex
Common sandpiper (Actitis hypoleucos) singing with tongue visible, Norway, June. - Ole  Jorgen Liodden
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Ole Jorgen Liodden / Common sandpiper (Actitis hypoleucos) singing with tongue visible, Norway, June.
Hippopotamus (Hippopotamus amphibius) rearing head out of water, creating spray, Lake Naivasha, Kenya. - Ole  Jorgen Liodden
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Ole Jorgen Liodden / Hippopotamus (Hippopotamus amphibius) rearing head out of water, creating spray, Lake Naivasha, Kenya.
Magnificent frigatebird (Fregata magnificens) male with gular pouch, Galapagos. - Ole  Jorgen Liodden
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Ole Jorgen Liodden / Magnificent frigatebird (Fregata magnificens) male with gular pouch, Galapagos.
Norway lemming (Lemmus lemmus) jumping aggressively, during the lemming population explosion, Vauldalen, Norway, May, 2011. - Erlend Haarberg
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Erlend Haarberg / Norway lemming (Lemmus lemmus) jumping aggressively, during the lemming population explosion, Vauldalen, Norway, May, 2011.
Arctic fox (Alopex lagopus) adult barking on rocks, Dovrefjell-Sunndalsfjella National Park, Norway, July. - Erlend Haarberg
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Erlend Haarberg / Arctic fox (Alopex lagopus) adult barking on rocks, Dovrefjell-Sunndalsfjella National Park, Norway, July.
Red deer (Cervus elaphus) stag calling in snowy mountains forest, portrait, Germany - Klein & Hubert
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Klein & Hubert / Red deer (Cervus elaphus) stag calling in snowy mountains forest, portrait, Germany
Group of King penguins (Aptenodytes patagonicus) walking on windy beach, one calling, Falklands. - Klein & Hubert
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Klein & Hubert / Group of King penguins (Aptenodytes patagonicus) walking on windy beach, one calling, Falklands.
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