Will Burrard-Lucas / Tufted gray langur (Semnopithecus priam) Close up portrait of head and shoulders. Minneriya National Park, Sri Lanka. September
Michael D. Kern / Kaleidoscope pattern formed from picture of Johnstons Chameleon (Chamaeleo johnstoni) face and legs. Restricted for Editorial use until December 2015
Nick Upton / Close-up of a Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Nick Upton / Male Thick-legged flower beetle (Oedemera nobilis), with a Pyramidal orchid (Anacamptis pyramidalis) pollinium attached to its head, nectaring on Common hogweed flowers (Heracleum sphondylium), Wiltsh...
Nick Upton / Two Dark green fritillary butterflies (Argynnis aglaja) and a Narrow-bordered five-spot burnet moth (Zygaena lonicerae) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland...
Nick Upton / Juvenile Goldenrod spider (Misumena vatia) camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, reacting to and catching a fly (Muscidae), Wiltshire, England, UK, June.
Espen Bergersen / Killer whale / orca (Orcinus orca). Spyhopping very close, showing eye. Kvanangen, Troms, Norway. November
John Chan / Flock of Western sandpipers (Calidris mauri) taking flight from salt marsh, Southern California, USA.
Ben Gillett / Slow motion clip of a female Appaloosa horse grooming a male, Bristol, England, UK, May.
Morley Read / Leaf frog (Agalychnis hulli) blinking its eyes and turning around, Orellana Province, Ecuador. (non-ex)
Morley Read / Developing embryo of the Ecuadorian monkey frog (Phyllomedusa ecuatoriana), showing heart beating and blood flowing, Morona Santiago Province, Ecuador, (non-ex)
Morley Read / Close-up of an Amazon leaf frog (Cruziohyla Craspedopus) tadpole developing in egg, Ecuador. (non-ex)
Ross Hoddinott / Common blue damselfly (Enallagma cyathigerum), close up portrait, Broxwater, Cornwall, UK. July.
Espen Bergersen / Aerial shot of a Humpback whale (Megaptera novaeangliae) swimming showing belly, Troms, Norway, January.
Michael Hutchinson / Marsh fritillary butterfly (Euphydryas aurinia) nectaring from a Buttercup (Ranunculus) before taking off, Devon, England, UK, June.
Michael Hutchinson / Close-up of a pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, showing tips of abdomens, Devon, England, UK, June.
Michael Hutchinson / Close-up of a pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, shot tilt downs, Devon, England, UK, June.
Jurgen Freund / Timelapse of a Cairns birdwing (Ornithoptera euphorion) butterfly caterpillar metamorphosing into a chrysalis, Queensland, Australia, April.
Jurgen Freund / Timelapse of a Cairns birdwing (Ornithoptera euphorion) butterfly caterpillar emerging from egg on an Dutchman's pipe (Aristolochia) leaf and feeding, Queensland, Australia, April.
Staffan Widstrand / Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) brooding chick aged 1-3 weeks, portrait. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick begging for food, on parent's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) egg hatching on father's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) pair mating. Atka Bay, Antarctica. May. Sequence 3/4.
Alex Hyde / Southern Elephant Seal bull (Mirounga leonina) close up portrait, King Haakon Bay, South Georgia. November.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
Georgette Douwma / Mantel of Spiny oyster (Spondylus varius). Indonesia. Indo-West Pacific.
Nick Upton / Red mason bees (Osmia bicornis) entering and leaving nest hole drilled into a log within an insect hotel, Gloucestershire, England, UK, April.
Mary McDonald / Adult Mountain gorilla (Gorilla beringei beringei) holding baby, Hirwa group, Volcanoes National Park, Rwanda
Paul Hobson / Spoonbill (Platalea leucorodia) preening, close up. Hungary. May.
Edwin Giesbers / Sword-grass moth (Xylena exsoleta) caterpillar on Field gladiolus (Gladiolus italicus). Cyprus. April.