David Tipling / Wandering albatross (Diomedea exulans) a pair displaying at dusk, Albatross Island, South Georgia January
David Tipling / Wandering albatross (Diomedea exulans) pair displaying at dusk in ritual bonding behaviour, Albatross Island in Bay of Isles, South Georgia January
David Tipling / Nightingale (Luscinia megarhynchos) singing, Camargue Provence France May
Markus Varesvuo / Japanese / Red-crowned crane (Grus japonicus) two calling and dancing, part of bonding and courtship display, Hokkaido Japan February
Tony Wu / Humpback whale (Megaptera novaeangliae) executing a dramatic tail slap, throwing up a cascade of water with its powerful fluke, Vava'u, Tonga, South Pacific
Tony Wu / Humpback whales (Megaptera novangliaea) unusual scene with two adults tail-slapping together in rough seas, Vava'u, Tonga, South Pacific
Tony Wu / Humpback whale (Megaptera novangliaea) executing an energetic tail slap, creating a cascade of seawater, Vava'u, Tonga, South Pacific
Alex Hyde / Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.
James Dunbar / Leaf cutter ants (Atta sp.) communicating. Guadeloupe National Park, Guadeloupe, Leeward Islands.
Klein & Hubert / Glaucous gull (Larus hyperboreus) standing on edge of ice calling, Svalbard, Norway
Bence Mate / Montezuma oropendolas (Psarocolius montezuma) perched either side of Brown-hooded parrot (Pyrilia haematotis) in rain, Laguna del Lagarto, Santa Rita, Costa Rica
Bence Mate / Black vulture (Coragyps atratus) face to face with King vulture (Sarcoramphus papa) Costa Rica.
Bence Mate / Orange chinned parakeets (Brotogeris jugularis) two looking at each other, Santa Rita, Costa Rica.
Tony Heald / African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana