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Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.  -  Markus Varesvuo
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Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.
African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...  -  Ben Cranke
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Ben Cranke / African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...
Blue jays (Cyanocitta cristata) confrontation in snow, April 2020. New York State, USA.  -  Marie Read
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Marie Read / Blue jays (Cyanocitta cristata) confrontation in snow, April 2020. New York State, USA.
Brown bear (Ursus arctos), two showing aggression at night. Finland. August.  -  Pal Hermansen
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Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.  -  Carol Walker
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Carol Walker / Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.
DUPLICATE Cuckoos (Cuculus canorus) squabbling, Kiskunsagi National Park, Pusztaszer, Hungary. February.  -  Bence  Mate
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Bence Mate / DUPLICATE Cuckoos (Cuculus canorus) squabbling, Kiskunsagi National Park, Pusztaszer, Hungary. February.
Great egrets (Ardea alba) fighting with necks intertwined,  Lake Csaj, Kiskunsagi National Park, Hungary. January.  -  Bence  Mate
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Bence Mate / Great egrets (Ardea alba) fighting with necks intertwined, Lake Csaj, Kiskunsagi National Park, Hungary. January.
Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.  -  Bence  Mate
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Bence Mate / Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.
Great egrets (Ardea alba) sky pointing, sizing each other up before fighting,  Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.  -  Bence  Mate
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Bence Mate / Great egrets (Ardea alba) sky pointing, sizing each other up before fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.
Two male Black grouse (Tetrao / Lyrurus tetrix) fighting, Kuusamo, Finland, April.  -  Markus Varesvuo
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Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting, Kuusamo, Finland, April.
Black capuchin, (Sapajus nigritus) female standing on hind legs, Rio de Janeiro, Brazil.  -  Mark  Bowler
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Mark Bowler / Black capuchin, (Sapajus nigritus) female standing on hind legs, Rio de Janeiro, Brazil.
German wasp (Vespula germanica) worker carrying rubbish, flying out of nest in air brick, Hertfordshire, England, UK.  July  -  Andy Sands
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Andy Sands / German wasp (Vespula germanica) worker carrying rubbish, flying out of nest in air brick, Hertfordshire, England, UK. July
Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.  -  Erlend Haarberg
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Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.
Bengal tiger (Panthera tigris tigris) 11 month cub, Ranthambhore National Park, India.  -  Andy  Rouse
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Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cub, Ranthambhore National Park, India.
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December.  -  Andy  Rouse
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Andy Rouse / Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December.
Scarlet Ibis (Eudocimus ruber) flying into roost sight at dusk, Hato La Aurora Reserve, Los Llanos, Colombia, South America.  -  Nick Garbutt
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Nick Garbutt / Scarlet Ibis (Eudocimus ruber) flying into roost sight at dusk, Hato La Aurora Reserve, Los Llanos, Colombia, South America.
Steller's Sea Eagle (Haliaeetus pelagicus) portrait, Hokkaido, Japan.  February.  -  Ben  Cranke
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Ben Cranke / Steller's Sea Eagle (Haliaeetus pelagicus) portrait, Hokkaido, Japan. February.
Bengal Tiger (Panthera tigris tigris) female 'Noor T39' with cubs in water. Ranthambore National Park, India.  -  Andy  Rouse
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Andy Rouse / Bengal Tiger (Panthera tigris tigris) female 'Noor T39' with cubs in water. Ranthambore National Park, India.
Hooded crow (Corvus coronix) and Eurasian Magpie (Pica pica) waiting to scavenging White tailed eagle's (Haliaeetus albicilla) food, Pusztazer, Hungary.  -  Bence  Mate
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Bence Mate / Hooded crow (Corvus coronix) and Eurasian Magpie (Pica pica) waiting to scavenging White tailed eagle's (Haliaeetus albicilla) food, Pusztazer, Hungary.
Lion (Panthera leo) cub aged about 2 years approaching with curiosity - remote wide angle perspective . Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.  -  Anup Shah
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Anup Shah / Lion (Panthera leo) cub aged about 2 years approaching with curiosity - remote wide angle perspective . Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America.  -  Edwin  Giesbers
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Edwin Giesbers / Female Jaguar (Panthera onca), captive, occurs in Southern and Central America.
Booby (Sula nebouxii) looking at camera underwater, Galapagos Island.  -  Michele  Westmorland
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Michele Westmorland / Booby (Sula nebouxii) looking at camera underwater, Galapagos Island.
Marbled poison dart frog (Epipedobates boulengeri) on leaf, Ecuador.  -  Bert  Willaert
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Bert Willaert / Marbled poison dart frog (Epipedobates boulengeri) on leaf, Ecuador.
South Herefordshire Huntsman confronting a Hunt Saboteur, Herefordshire, UK  -  Tom  Gilks
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Tom Gilks / South Herefordshire Huntsman confronting a Hunt Saboteur, Herefordshire, UK
Lion-tailed macaque (Macaca silenus) female portrait. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India. Apr 2013.  -  Fiona Rogers
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Fiona Rogers / Lion-tailed macaque (Macaca silenus) female portrait. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India. Apr 2013.
Grizzly bear (Ursus arctos horribilis) portrait, Lake Clark National Park, Alaska, USA, September.  -  Ingo Arndt
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Ingo Arndt / Grizzly bear (Ursus arctos horribilis) portrait, Lake Clark National Park, Alaska, USA, September.
African penguins (Spheniscus demersus) diving underwater, False Bay, Cape Town, South Africa.  -  Chris & Monique Fallows
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Chris & Monique Fallows / African penguins (Spheniscus demersus) diving underwater, False Bay, Cape Town, South Africa.
Wolf (Canis lupus) approaches a small herd of Red deer (Cervus elephas) in snow, Abruzzo, Italy. Highly commended in the Mammals category of the GDT Competition 2013  -  Bruno DAmicis,Bruno  D'Amicis
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Bruno DAmicis,Bruno D'Amicis / Wolf (Canis lupus) approaches a small herd of Red deer (Cervus elephas) in snow, Abruzzo, Italy. Highly commended in the Mammals category of the GDT Competition 2013
Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...  -  Bence  Mate
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Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...
Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Nature's Best Photography Windland Smith Rice International Awards Competition...  -  David  Pattyn
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David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Nature's Best Photography Windland Smith Rice International Awards Competition...
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February  -  David Pattyn
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David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa.  -  Tony Phelps
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Tony Phelps / Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa.
African Lioness (Panthera leo) portrait, with aggressive expression because of cubs nearby Maasai Mara, Kenya, Africa  -  Andy Rouse
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Andy Rouse / African Lioness (Panthera leo) portrait, with aggressive expression because of cubs nearby Maasai Mara, Kenya, Africa
Portrait of a spotted sunfish (Lepomis punctatus) in front of reeds in a river. Rainbow River, Florida, United States of America.  -  Alex  Mustard
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Alex Mustard / Portrait of a spotted sunfish (Lepomis punctatus) in front of reeds in a river. Rainbow River, Florida, United States of America.
Two male Black grouse (Lyrurus tetrix) displaying at lek, Kuusamo, Finland, April.  -  Markus Varesvuo,Markus Varesvuo
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Markus Varesvuo,Markus Varesvuo / Two male Black grouse (Lyrurus tetrix) displaying at lek, Kuusamo, Finland, April.
Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July  -  Anup Shah,Anup Shah
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Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July
Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya  -  Anup Shah,Anup Shah
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Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya
Indri (Indri indri) portrait in tropical rainforest habitat. Madagascar.  -  Andy  Rouse
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Andy Rouse / Indri (Indri indri) portrait in tropical rainforest habitat. Madagascar.
African Elephant (Loxodonta africana) in defensive posture towards safari vehicle, Sabi Sands Private Game Reserve, South Africa  -  Suzi Eszterhas
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Suzi Eszterhas / African Elephant (Loxodonta africana) in defensive posture towards safari vehicle, Sabi Sands Private Game Reserve, South Africa
African Lion (Panthera leo) fending off Cape Buffalo (Syncerus caffer), Africa  -  Pete Oxford
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Pete Oxford / African Lion (Panthera leo) fending off Cape Buffalo (Syncerus caffer), Africa
African Lion (Panthera leo) group killing Cape Buffalo (Syncerus caffer) while fending off rest of herd, Africa  -  Pete Oxford
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Pete Oxford / African Lion (Panthera leo) group killing Cape Buffalo (Syncerus caffer) while fending off rest of herd, Africa
Black-backed Jackal (Canis mesomelas) confronting a Cape Vulture (Gyps coprotheres), Giant's Castle Nature Reserve, Drakensberg, South Africa  -  Winfried Wisniewski
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Winfried Wisniewski / Black-backed Jackal (Canis mesomelas) confronting a Cape Vulture (Gyps coprotheres), Giant's Castle Nature Reserve, Drakensberg, South Africa
Northern Gannet (Morus bassanus) defending partner, Saltee Island, Ireland  -  Jasper Doest
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Jasper Doest / Northern Gannet (Morus bassanus) defending partner, Saltee Island, Ireland
Northern Gannet (Morus bassanus) pair fighting, Saltee Island, Ireland  -  Jasper Doest
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Jasper Doest / Northern Gannet (Morus bassanus) pair fighting, Saltee Island, Ireland
Horned Anole (Anolis proboscis) males confronting on branch, Mindo, Pichincha, Ecuador. Sequence 7 of 10  -  James Christensen
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James Christensen / Horned Anole (Anolis proboscis) males confronting on branch, Mindo, Pichincha, Ecuador. Sequence 7 of 10
Horned Anole (Anolis proboscis) males confronting on branch, Mindo, Pichincha, Ecuador. Sequence 3 of 10  -  James Christensen
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James Christensen / Horned Anole (Anolis proboscis) males confronting on branch, Mindo, Pichincha, Ecuador. Sequence 3 of 10
Golden Snub-nosed Monkey (Rhinopithecus roxellana) male with juvenile in the snow running towards another on a rock, Qinling Mountains, China  -  Stephen Belcher
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Stephen Belcher / Golden Snub-nosed Monkey (Rhinopithecus roxellana) male with juvenile in the snow running towards another on a rock, Qinling Mountains, China
Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Kuusamo, Finland, April.  -  Markus Varesvuo
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Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Kuusamo, Finland, April.
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