Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting at lek. Utajarvi, Finland, April.
Ben Cranke / African fish eagle (Haliaeetus vocifer) pressurises a Saddle-billed stork (Ephippiorhynchus senegalensis) into dropping a freshly caught fish, so that it can be stolen for a meal. Liuwa Plain National...
Marie Read / Blue jays (Cyanocitta cristata) confrontation in snow, April 2020. New York State, USA.
Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
Carol Walker / Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA.
Bence Mate / DUPLICATE Cuckoos (Cuculus canorus) squabbling, Kiskunsagi National Park, Pusztaszer, Hungary. February.
Bence Mate / Great egrets (Ardea alba) fighting with necks intertwined, Lake Csaj, Kiskunsagi National Park, Hungary. January.
Bence Mate / Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.
Bence Mate / Great egrets (Ardea alba) sky pointing, sizing each other up before fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January.
Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting, Kuusamo, Finland, April.
Mark Bowler / Black capuchin, (Sapajus nigritus) female standing on hind legs, Rio de Janeiro, Brazil.
Andy Sands / German wasp (Vespula germanica) worker carrying rubbish, flying out of nest in air brick, Hertfordshire, England, UK. July
Erlend Haarberg / Arctic fox (Alopex lagopus) cubs looking curiously, Dovrefjell-Sunndalsfjella National Park, Norway, July.
Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cub, Ranthambhore National Park, India.
Andy Rouse / Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December.
Nick Garbutt / Scarlet Ibis (Eudocimus ruber) flying into roost sight at dusk, Hato La Aurora Reserve, Los Llanos, Colombia, South America.
Tom Gilks / South Herefordshire Huntsman confronting a Hunt Saboteur, Herefordshire, UK
Fiona Rogers / Lion-tailed macaque (Macaca silenus) female portrait. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India. Apr 2013.
Ingo Arndt / Grizzly bear (Ursus arctos horribilis) portrait, Lake Clark National Park, Alaska, USA, September.
Chris & Monique Fallows / African penguins (Spheniscus demersus) diving underwater, False Bay, Cape Town, South Africa.
Bruno DAmicis,Bruno D'Amicis / Wolf (Canis lupus) approaches a small herd of Red deer (Cervus elephas) in snow, Abruzzo, Italy. Highly commended in the Mammals category of the GDT Competition 2013
Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...
David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Nature's Best Photography Windland Smith Rice International Awards Competition...
David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Tony Phelps / Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa.
Andy Rouse / African Lioness (Panthera leo) portrait, with aggressive expression because of cubs nearby Maasai Mara, Kenya, Africa
Alex Mustard / Portrait of a spotted sunfish (Lepomis punctatus) in front of reeds in a river. Rainbow River, Florida, United States of America.
Markus Varesvuo,Markus Varesvuo / Two male Black grouse (Lyrurus tetrix) displaying at lek, Kuusamo, Finland, April.
Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July
Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya
Andy Rouse / Indri (Indri indri) portrait in tropical rainforest habitat. Madagascar.
Suzi Eszterhas / African Elephant (Loxodonta africana) in defensive posture towards safari vehicle, Sabi Sands Private Game Reserve, South Africa
Pete Oxford / African Lion (Panthera leo) fending off Cape Buffalo (Syncerus caffer), Africa
Pete Oxford / African Lion (Panthera leo) group killing Cape Buffalo (Syncerus caffer) while fending off rest of herd, Africa
Winfried Wisniewski / Black-backed Jackal (Canis mesomelas) confronting a Cape Vulture (Gyps coprotheres), Giant's Castle Nature Reserve, Drakensberg, South Africa