Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Nature Production / Bombardier Beetle (Pheropsophus jessoensis) protecting itself by ejecting a boiling, noxious chemical spray, Nagasaki, Japan.

Ole Jorgen Liodden / Arctic tern (Sterna paradisaea) attacking an Arctic Fox (Vulpes / Alopex lagopus) protecting its nest site, Spitsbergen, Svalbard, Norway. Highly commended, MAMMALS, GDT 2011 competition

Anup Shah / African fish eagle {Haliaeetus vocifer} defends Lesser flamingo carcass from Marabou storks {Leptoptilos crumeniferus}, Lake Nakuru, Kenya.

Anup Shah / African fish eagle {Haliaeetus vocifer} fighting Marabou storks {Leptoptilos crumeniferus} over Lesser flamingo carcass, Lake Nakuru, Kenya.

Anup Shah / Juvenile Tawny or African Sea Eagle defending flamingo carcass from Marabou stork {Lepitoptilos crumeniferous}

Doug Perrine / Great white shark jaws open at surface {Carcharodon carcharias} South Africa (Non-ex).

Brandon Cole / Great white shark (Carcharodon carcharias) attacking below surface. Dyer Is, South Africa NOT FOR SALE IN USA, DIGITALLY ENHANCED

Owen Newman / Male Red kangaroos fighting {Macropus rufus} Sturt NP, New South Wales Australia

Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month old cubs play fighting, Ranthambhore National Park, India.

Erlend Haarberg / Black-legged kittiwakes (Rissa tridactyla) fighting mid air, Hornoeya. Finnmark, Norway, July.

Anup Shah / Lioness (Panthera leo) rejecting a male's advances Maasai Mara National Reserve, Kenya.

David Pattyn / Great crested grebe (Podiceps cristatus) attacking another grebe in the mating season during a territorial dispute. The Netherlands. April.

David Pattyn / Shelduck (Tadorna tadorna) male trying to grab a female to mate with her. The Netherlands. April.

David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. April.

David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. March 2013

David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. The Netherlands. April 2014

David Pattyn / Great crested grebe (Podiceps cristatus) adults fighting in a territorial dispute in the mating season. The Netherlands.april 2014

David Pattyn / Black necked grebe (Podiceps nigricollis) pair chasing away another grebe during the courtship season. The Netherlands.April 2014

David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. The Netherlands. April.

David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. April.

Erlend Haarberg / Mountain hares (Lepus timidus) boxing, Vauldalen, Sor-Trondelag, Norway, May.

Erlend Haarberg / Mountain hares (Lepus timidus) boxing, Vauldalen, Sor-Trondelag, Norway, May.

Alex Hyde / Baboon Spider (Pterinochilus murinus) in aggressive posture, photographed on a white background. Captive, originating from Africa.

Eric Medard / Ibex (Capra ibex) males fighting, Jura, Switzerland, captive

Jabruson Motion / Two male African forest elephants (Loxodonta cyclotis) play fighting in a forest clearing, Dzanga Bai, Dzanga-Ndoki National Park, Sangha-Mbaere Prefecture, Central African Republic.

Jabruson Motion / Two male African forest elephants (Loxodonta cyclotis) play fighting in a forest clearing, Dzanga Bai, Dzanga-Ndoki National Park, Sangha-Mbaere Prefecture, Central African Republic.

Pål Hermansen,Pal Hermansen / Song thrush (Turdus philomelos) with grub prey on the dashboard of an abandoned car in 'car graveyard' Varmland, Sweden, July

Pål Hermansen,Pal Hermansen / Nuthatch (Sitta europaea) chasing away and Black tit (Periparus ater) in car graveyard, Varmland, Sweden, December

Pal Hermansen / Sparrowhawk (Accipiter nisus) attacking Jay (Garrulus glandarius) Telemark, Norway, September

Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...

Jabruson / Hippopotamuses fighting (Hippopotamus amphibius) taken before major crash in numbers in the 1990s due to political instability and hunting. Rwindi sector of the Virunga National Park. North Eastern Za...

Konrad Wothe / Aggressive Mute swan (Cygnus olor) Upper Bavaria, Germany, Europe

Denis-Huot,Denis-Huot / Lions (Panthera leo) mating, Masai-Mara Game Reserve, Kenya

Visuals Unlimited / Grizzly Bear (Ursus arctos horribilis) roaring head profile, Montana, USA. Captive.

Doug Perrine / Hawaiian monk seals (Monachus schauinslandi) aggressive male attacks a five and a half week pup, mother (right) approaches to intervene; Papaloa Beach, Kalaupapa, Molokai, Hawaii, USA

Loic Poidevin,Loic Poidevin / Dalmatian pelicans (Pelecanus crispus) squabbling over fish, Lake Kerkini, Greece, February

Markus Varesvuo,Markus Varesvuo / Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April.

Markus Varesvuo,Markus Varesvuo / Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April.

Markus Varesvuo,Markus Varesvuo / Two male Black grouse (Lyrurus tetrix) displaying at lek, Kuusamo, Finland, April.

Markus Varesvuo,Markus Varesvuo / Black Grouse (Lyrurus tetrix) Kuusamo Finland April 2013

Steven Kazlowski / Polar bear (Ursus maritimus) 3-year-olds play fighting on newly formed pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer). Masai Mara National Reserve, Kenya, July

Anup Shah,Anup Shah / Male lion (Panthera leo) in confrontation with Cape buffalo (Syncerus caffer) Masai Mara National Reserve, Kenya

Denis-Huot,Denis-Huot / Black rhino (Diceros bicornis) looking threatening, Nakuru National Park, Kenya

Denis-Huot,Denis-Huot / Lion (Panthera leo) mating, Masai-Mara Game Reserve, Kenya. Vulnerable species.

Igor Shpilenok,Igor Shpilenok / Red Foxes (Vulpes vulpes) fighting outside ranger cabin window. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, February.