Andrew Peacock / Minden / Taiga and snow-covered mountains in autumn, Glacier Bay National Park, Alaska
Duncan Usher / Autumn forest
Silvia Reiche / Oldworld Swallowtail (Papilio machaon) butterfly on red autumn leaf, Hoogeloon, Noord-Brabant, Netherlands
Heike Odermatt / Caribou (Rangifer tarandus) mother with calf, Flatruet, Sweden
Guy Edwardes / Magpie inkcap fungus (Coprinopsis picacea). New Forest National Park, England, UK. October.
Mark MacEwen / Angel Falls, the world's highest uninterrupted waterfall with a fall of 807m, flowing from Auyan-tepui, a table-top mountain. Helicopter used in filming at base of waterfall with people beside it....
Ben Hall / Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October.
Tony Heald / Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October.
Edwin Giesbers / Three Long-eared owls (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Birds in their Habitat Category winner, Expos Photos Aves Emotion'Ailes Competition 2020.
Edwin Giesbers / Long-eared owl (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Second Prize Winner, Birds in their Environment Category, Bird Photographer of the Year (BPOTY) 2019...
Marie Read / Blue jay (Cyanocitta cristata) in flight carrying an acorn that it will cache for winter food supply, New York, USA, October.
Danny Green / Wolf (Canis lupus) standing at woodland edge. Finland, April.
Guy Edwardes / Autumn colour around the Falls of Clyde, Lanarkshire, Scotland, UK, October 2006.
Jack Dykinga / Arizona sycamore (Platanus wrightii) roots and leaves amongst pebbles. In riparian area, Chiricahua Mountains, Coronado National Forest, Arizona, USA. November.
Guy Edwardes / Corfe Castle in early morning mist, Dorset, England, UK, October 2018.
Andres M. Dominguez / Crested tit (Lophophanes cristatus) Sierra de Grazalema Natural Park. Southern Spain. September.
Andres M. Dominguez / Cork trees (Quercus suber) stripped of bark, in forest. Los Alcornocales Natural Park, Cortes de la Frontera, southern Spain, November.
Andres M. Dominguez / Hawfinch (Coccothraustes coccothraustes) perched on branch amongst Hawthorn (Crataegus sp) berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February.
Ingo Arndt / Aerial shot over a forest of Southern beech trees (Nothofagus) in autumn, Torres del Paine National Park, Patagonia, Chile, April, 2018.
Jack Dykinga / Rainbow over Long Canyon with fall coloured Cottonwood (Populus) trees against canyon walls, Burr Trail, Grand Staircase-Escalante National Monument, Utah, USA, October 2018.
Jack Dykinga / Waterfall over sandstone with cottonwood tree, Long Canyon , Burr Trail, Grand Staircase-Escalante National Monument, Utah, USA. October.
Jack Dykinga / Morning light with autumn coloured Cottonwoods (Populus sp) trees lining the river. Valley of the Gods, Colorado Plateau, of the Great Basin Desert, Utah, USA, November 2018.
Nick Hawkins / Chum salmon (Oncorhynchus keta) migrate up a small river near Bella Bella, British Columbia, Canada. September
Nick Hawkins / Atlantic salmon (Salmon salar) under river ice. Quebec, Canada. October.
Marie Read / Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October.
Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.
Marie Read / Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January.
Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September
Bruno D'Amicis / Red fox (Vulpes vulpes) walking along fallen trunk of old Beech (Fagus sylvatica) tree, Coppo del Principe old-growth beech forest in autumn. Abruzzo, Lazio and Molise National Park / Parco Nazionale...
Sven Zacek / Bog surrounding river in morning light. Emajoe Suursoo Nature Reserve, Tartumaa, Estonia. October 2015.
Sven Zacek / Flooded meanders of Halliste River, aerial view. Soomaa National Park, Parnumaa, Western Estonia. October 2017.
Philippe Clement / Two Lynx (Lynx lynx) kittens play fighting in front of den entrance, Bavarian Forest National Park, Germany, October. Captive.
TJ Rich / Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
David Allemand / Birch tree (Betula sp) next to waterfall, Iceland. Runner-Up in the Plants category of the International OasisPhotoContest 2018
Erlend Haarberg / Red squirrel (Sciurus vulgaris) hiding in a tree trunk, Klaebu, Norway, September.
Guy Edwardes / View of hikers from behind the Pericnik Waterfall, Slovenia, October 2018.
Bernard Castelein / Beech tree (Fagus sylvatica) showing roots, Peerdsbos, Brasschaat, Belgium