John Waters / Honey bee (Apis mellifera) approaching hive entrance, with trails of departing bees visible. Urban garden, Bristol, UK, May.

John Waters / Honey bee (Apis mellifera) flying close to hive entrance, with flight trails visible. Urban garden, Bristol, UK, May.

Bence Mate / African buffalo (Syncerus caffer) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / Hippopotamus (Hippopotamus amphibius) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / African buffalo (Syncerus caffer) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017.

Bence Mate / Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night. Winner of the Mammals Category of the GDT Awards 20...

Cheryl-Samantha Owen / West coast rock lobster (Jasus lalandii) commercial fishing boat (James Archer) leaves the Saldhana Bay harbour before the break of dawn. The lights of the town and the lighthouse flash on the horizon...

John Abbott / Composite image of flying insects visiting blacklight, Texas, USA, September

Bence Mate / Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition.

Charlie Hamilton James / Kingfishers (Alcedo atthis) adult male diving, strobed flash, Halcyon River, England

Stephen Dalton / Blue tit {Parus caerulus} flight sequence, multiflash image, UK

Stephen Dalton / Meadow grasshopper {Chorthippus parellelus} jumping sequence, multiflash, UK

Stephen Dalton / Pair of Common pipistrelle bats {Pipistrellus pipistrellus} in flight, UK

Stephen Dalton / Greater horseshoe bat {Rhinolophus ferrumequinum} flight sequence, multiflash image, UK

Stephen Dalton / Green lacewing {Chrysopa sp} flight sequence, multiflash image, UK

Stephen Dalton / Cat flea {Ctenocephalides felis} leaping sequence, multiflash image

Stephen Dalton / Ringneck / Java dove {Streptopelia roseogrisea} flight sequence, multiflash image, UK

Stephen Dalton / Ringneck / Java dove {Streptopelia roseogrisea} flight sequence, multiflash image, controlled conditions

Stephen Dalton / Domestic cat {Felis catus} leaping sequence, multiflash image

Stephen Dalton / Common european tree frog {Hyla arborea} jumping sequence, multiflash image, controlled conditions, UK

Stephen Dalton / Common / European treefrog {Hyla arborea} jumping sequence, multiflash image, controlled conditions

Stephen Dalton / Common / European treefrog {Hyla arborea} jumping sequence, multiflash image, controlled conditions

Stephen Dalton / Common frog (Rana temporaria) jumping sequence, multiflash image, UK, controlled conditions

Laurie Campbell / Flash flood, Kinloch Rannoch, Perthshire, Scotland, UK, June

Jack Dykinga / Sand dunes with cracked clay marking the last flash flood, Creosote bushes (Larrea tridentata) flowering in the background, Mosquito Flats, Death Valley National Park, California, USA

Christophe Courteau / Small Spotted Genet (Genetta genetta) poking head out of a tree at night, Botswana

Jack Dykinga / Water from flash floods cascades down Marble Canyon into the Colorado River, Grand Canyon NP, Arizona, USA

Doug Gimesy / Julie Jennings, senior wildlife carer for Port Stephens Koalas, move bushfire a burnt Koala (Phascolarctos cinereus) named 'Flash' from his enclosure to allow her to clean the enclosure and check and...

Doug Gimesy / Julie Jennings, senior wildlife carer for Port Stephens Koalas, feeds bushfire burnt koala (Phascolarctos cinereus) named 'Flash', a supplementary feed, which is in addition to the eucalyptus leave Fl...

Doug Gimesy / Julie Jennings, senior wildlife carer for Port Stephens Koalas, feeds bushfire burnt koala (Phascolarctos cinereus) named 'Flash', a supplementary feed, which is in addition to the eucalyptus leave Fl...

Doug Gimesy / Portrait of a burnt koala (Phascolarctos cinereus) named 'Flash', a victim of the bushfires at Hillville near Taree (NSW). He arrived into care with a burnt nose, burnt hands and feet. Despite...

Doug Gimesy / Bushfire victim Koala (Phascolarctos cinereus) named 'Flash' in his enclosure eating eucalyptus leaves. Flash was caught in a bushfire at Hillville near Taree (NSW) in November, 2019. He arrived to ca...

Doug Gimesy / Tourists photographing Little penguin (Eudyptula minor) with flash photography, at night. St Kilda breakwater, St Kilda Pier, Melbourne, Victoria, Australia. June 2016.

SCOTLAND: The Big Picture / Firework anemone (Pachycerianthus multiplicatus) at night, taken with flash, Loch Duich, Scotland, UK, January.

SCOTLAND: The Big Picture / Phosphorescent sea pen (Pennatula phosphorea) at night, taken with flash, Loch Duich, Scotland, UK, January.

Paul Hobson / Pyramidal Orchid (Anacamptis pyramidalis), three flowers with railway viaduct in background. Sprotborough Flash, South Yorkshire, England, UK. June.

Ashley Cooper / Gentoo penguin (Pygoscelis papua) with huge wound, probably from Leopard seal (Hydrurga leptonyx) attack, Hannah Point, Livingstone Island, South Shetland Islands, Antarctica.

Ashley Cooper / Gentoo penguin (Pygoscelis papua) with huge gash, probably from Leopard seal (Hydrurga leptonyx) attack, Hannah Point, Livingstone Island, South Shetland Islands, Antarctica.

Will Watson / Natural oil released from dead Copepods under anoxic conditions on a back marsh, Upton Warren flash pool, Worcestershire, England, August 2017.

Will Watson / Natural oil released from dead Copepods under anoxic conditions on a back marsh, Upton Warren flash pool, Worcestershire, England, UK, August.

Will Watson / Anoxic mud with Cynobacteria and rivulets of black anoxic bacteria on the edge of Upton Warren saline flash pool, Worcestershire, England, UK, August.

John Abbott / Cicindela sexguttata (Tiger Beetle) flying image captured using high speed flash and laser triggered camera. Bastrop County, Texas, USA. Controlled conditions. March

John Abbott / Evening bat (Nycticeius humeralis) flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July

John Abbott / Cave myotis bat (Myotis velifer) flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July

John Abbott / Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July

John Abbott / Burying beetle (Nicrophorus orbicollis) flying shots taken with high speed flash. Lamar County, Texas, USA Controlled conditions. June

Solvin Zankl / Eye-flash squid (Abralia veranyi) which has 550 light producing photophores to help reduce the squids outline, Atlantic Ocean off Cape Verde. Captive.

Solvin Zankl / Eye-flash squid (Abralia veranyi) from Atlantic Ocean off Cape Verde. Captive.