Chien Lee / Minden / Bornean Smaller Flying Frog (Rhacophorus borneensis), Kubah National Park, Sarawak, Malaysia
Chien Lee / Minden / Black-headed Cat Snake (Boiga nigriceps) swallowing alive Harlequin Flying Tree Frog (Rhacophorus pardalis) prey, Kubah National Park, Sarawak, Malaysia, sequence 1 of 4
Jan Wegener / Common Nighthawk (Chordeiles minor) male in display flight, Winnie, Texas
Tony Wu / Japanese dwarf flying squirrel (Pteromys volans orii) gliding through snowy forest, Hokkaido, Japan.
Stephen Dalton / Indian leaf butterfly (Kallima paralekta) in flight.
Stephen Dalton / Barn owl (Tyto alba) in flight approaching bell-tower, carrying rodent prey to young in nest. Sussex, UK.
Oscar Dewhurst / Canada geese (Branta canadensis) in flight low over lake at sunrise in mist. London, UK. November
Stephen Dalton / Orange tip butterfly (Anthocharis cardamines) male in flight amongst Lady's smock or Cuckooflower (Cardamine pratensis),England.
Stephen Dalton / Black witch moth (Ascalapha odorata) in flight, Venezuela. This large moth has a 7 inch wingspan. Wingscales being shed during flight.
Sven Zacek / Brimstone butterfly(Gonepteryx rhamni) male and female during mating flight, early spring. Tartumaa county, Southern Estonia. May.
David Tipling / Red grouse (Lagopus lagopus) male in flight over Heather. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.
David Tipling / Red grouse (Lagopus lagopus) male in flight. Grinton Moor, above Swaledale, Yorkshire Dales National Park, England, UK. July.
David Tipling / Flock of Knot (Calidris canutus) chased by a Peregrine as they arrive to roost at high tide, Cormorants in foreground, Snettisham RSPB Reserve, The Wash, Norfolk, September.
Andres M. Dominguez / Crested tit (Lophophanes cristatus) about to land on Elmleaf blackberry (Rubus ulmifolius) plant, Sierra de Grazalema Natural Park, Southern Spain, January.
Jeff Foott / California condor (Gymnogyps californianus) landing, forest in background. Bird with tags on wings. Near San Pedro Martir National Park, Northern Baja California, Mexico. 2017.
Pal Hermansen / Whooping swan (Cygnus cygnus) in flight, blurred image. Tysslingen, Orebro, Sweden. March.
Pal Hermansen / Fishing boat surrounded by flock of Gulls, blurred image. Troms and Finnmark, Norway. November 2014.
Pal Hermansen / Oystercatcher (Haematopus ostralegus) landing on coast in midnight sun. Lofoten, Norway. July.
Neil Aldridge / A lesser horseshoe bat (Rhinolophus hipposideros) flying over grassland habitat at night. Devon, UK.
Eric Medard / Hen harrier (Circus cyaneus) flying / hovering in the rain, Mayenne, France. April.
Markus Varesvuo / Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February.
Lucas Bustamante / Monarch butterflies (Danaus plexippus) flying during a warm morning. When the overwintering period draws to an end large numbers become active. Michoacan, Mexico, February.
John Waters / Honey bee (Apis mellifera) approaching hive entrance, with trails of departing bees visible. Urban garden, Bristol, UK, May.
John Waters / Honey bee (Apis mellifera) flying close to hive entrance, with flight trails visible. Urban garden, Bristol, UK, May.
Guy Edwardes / Long-billed curlew (Numenius americanus) taking off along shoreline. Fort De Soto Park, Florida, USA. April.
Guy Edwardes / Blue tit (Cyanistes caeruleus) in flight. Slovenia. February.
Tony Wu / Japanese dwarf flying squirrel (Pteromys volans orii) male sitting in tree, portrait. Hokkaido, Japan. March
Staffan Widstrand / Great white egret (Egretta / Ardea alba) Pusztaszer protected landscape, Kiskunsagi, Hungary, May
Philippe Clement / Blue tit (Cyanistes caeruleus) collecting cotton wool for nest building before flying out of frame, Belgium, April.
Marie Read / Blue jay (Cyanocitta cristata) in flight carrying an acorn that it will cache for winter food supply, New York, USA, October.
George Sanker / Blue jay (Cyanocitta cristata) flying, head on, Acadia National Park, Maine, USA. May.
James Dunbar / Beefly (Bombyliidae sp.) feeding at flower, April, UK
Simon Littlejohn / Juvenile Lammergeier vulture (Gypaetus barbatus) in flight, mobbed by a raven, Grand Bargy cirque, Haute Savoie, France, October.
Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male, before flying off in response to a female, Bedfordshire, England, UK, May.
Morley Read / Slow motion shot of a Rufous tailed hummingbird (Amazilia tzacatl) drinking nectar from a Heliconia flower, Mindo, Ecuador.
Morley Read / Slow motion shot of a Violet tailed sylph hummingbird (Aglaiocercus coelestis) drinking nectar from a Heliconia flower, Mindo, Ecuador.
Markus Varesvuo / White-tailed eagle (Haliaeetus albicilla) with backlit wings, and water spray, Norway, October
Nick Upton / Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December.
Ben Cranke / African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia
Jack Dykinga / Yellow-headed blackbird (Xanthocephalus xanthocephalus) flock murmuration as they take off from the marsh at dawn, Whitewater Draw, Arizona State Game and Fish Reserve, USA. January.
Sven Zacek / Ural Owl (strix uralensis) female with talons ready to attack a ringing ornithologist, Tartumaa county, Southern Estonia. May.
Nick Upton / Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire,...
Michael Hutchinson / Wide angle shot of a female Goldenrod spider (Misumena vatia) on Annual honesty (Lunaria annua) flower, with flies flying nearby, Bristol, England, UK, April.
Guy Edwardes / Orange nectar bat (Lonchophylla robusta) feeding on a banana flower, lowland rainforest, Costa Rica. November.
Guy Edwardes / Two Pallas's long-tongued bats (Glossophaga soricina) feeding from banana flower, lowland rainforest, Costa Rica. November.
Guy Edwardes / Orange Nectar Bat (Lonchophylla robusta), lowland rainforest, Costa Rica. November.
Guy Edwardes / Pallas's long-tongued Bat (Glossophaga soricina) feeding from Flowering banana (Musa ornata), Lowland rainforest, Costa Rica. November.
Guy Edwardes / Pallas's long-tongued Bat (Glossophaga soricina) feeding from Calabash gourd flower (Crescentia cujete) , lowland rainforest, Costa Rica. November.