John Cancalosi / Sonoran desert toad (Incilius alvarius) pair mating, with trail of spawn in water. Arizona, USA.

Morley Read / Slow motion clip of a South American common toad (Rhinella margaritifera) leaping, Orellana Province, Ecuadorian Amazon.

Morley Read / Close-up portrait of a South American common toad (Rhinella margaritifera) amongst leaf litter, Orellana Province, Ecuadorian Amazon.

Morley Read / Slow motion clip of a Guacamayo plump toad (Osornophryne guacamayo) walking, Orellana Province, Ecuador.

Morley Read / Rio Faisanes stubfoot toad (Atelopus coynei), Carchi Province, Ecuador, 2017. Critically endangered.

John Cancalosi / American toads (Anaxyrus americanus), several males competing to mate with female, Maryland, USA, April.

Kerstin Hinze / Male Common european toad (Bufo bufo) in amplexus with a dead fish, Lower Saxony, Germany, April.

John Waters / Male Common European toad (Bufo bufo) approaching female at surface, Priddy, Somerset, England, UK, April.

John Waters / Wide-angle shot of a pair of Common European toads (Bufo bufo) in amplexus, jumping into a pond, Priddy, Somerset, England, UK, April.

Remi Masson / Common toads (Bufo bufo) mating surrounded by strings of toadspawn. Ain, Alps, France. First prize in the category "The Underwater World, GDT European Wildlife Photographer of the Year 2014 competiti...

Lucas Bustamante / Rio pescado harlequin toad (Atelopus balios) pair on branch, female on the left, Azuay, Ecuador, Critically endangered species, possibly extinct.

Remi Masson / Male Common toads (Bufo bufo) fighting underwater during mating season, with a pair mating Ain, Rhone-Alps, France, March.

John Cancalosi / Male American toad (Bufo americanus) calling to attract female, Ithaca, New York, USA, May.

MYN / Paul van Hoof / Common spadefoot toad (Pelobates fuscus) composite showing metamorphosis from larva to adult, The Netherlands, meetyourneighbours.net project

Sam Hobson / Common toads (Bufo bufo) in amplexus, Bristol, UK, March.

Alastair MacEwen / Time-lapse of toadspawn from a Common toad (Bufo bufo) developing, emerging and feeding on nutrient jelly, Somerset, England, UK, April.

John Waters / Underwater view of a group of Common toads (Bufo bufo) in a mating ball in a pond, Priddy, Somerset, England, UK, March.

John Waters / Underwater view of a group of Common toads (Bufo bufo) spawning in a pond, some in amplexus, with spawn, Priddy, Somerset, England, UK, March.

John Waters / Group of Common toads (Bufo bufo) in a mating ball in a pond, Priddy, Somerset, England, UK, March.

John Waters / Group of Common toads (Bufo bufo) in a mating ball in a pond, Priddy, Somerset, England, UK, March.

Benjamin Barthelemy / Pair of Common european toads (Bufo bufo) pair in amplexus in the Lac des Corbeaux, with frogspawn, Ballons des Vosges Regional Natural Park, Vosges, Lorraine, France, April

Bert Willaert / Juvenile South American Common / Crested Forest Toad (Rhinella margaritifera 'complex'). Ecuador, South America.

Daniel Heuclin / Portrait of a Crested Forest Toad (Rhinella margaritifer). Controlled conditions. French Guyana, August.

Miles Barton / Panamanian Golden Frog (Atelopus ziteki) Panama, Featured in BBC NHU series 'Life in Cold Blood'. Species now extinct in the wild

Miles Barton / Panamanian golden frog (Atelopus ziteki) reacting to the image of a frog on a video monitor. Experiment carried out to reveal the purpose of the frog's waving behaviour - to attract mates and deter ri...

Tim Laman / Frog {Rana similis} Sierra Madre National Park, Luzon, Philippines

Tim MacMillan / John Downer Pr / Multiple exposure study of various frogs gliding. Those with more webbing between their toes glide further (Resolution restriction - image digitised from film, 'Weird Nature' tv series)

Tim MacMillan / John Downer Pr / Multiple exposure study of various frogs shows that those with more foot webbing glide further (Resolution restriction - image digitised from film, 'Weird Nature' tv series)

Pete Oxford / Crested toads on leaf litter {Bufo typhonius}, Amazon, Ecuador Captive

MYN / Seth Patterson / Plains Spadefoot Toad (Spea bombifrons) male portrait, Hidalgo County, Lower Rio Grande Valley, Texas, United States of America, North America, September. Meetyourneighbours.net project

MYN / Seth Patterson / Plains Spadefoot Toad (Spea bombifrons) male portrait, Hidalgo County, Lower Rio Grande Valley, Texas, United States of America, North America, September. Meetyourneighbours.net project

Emanuele Biggi / Golden frog (Atelopus varius) in captivity at the El Valle de Anton Conservation Centre (EVACC), Panama.

Emanuele Biggi / Harlequin frog (Atelopus varius) mating couple in captivity at the El Valle de Anton Conservation Centre (EVACC), critically endangered, Panama

Emanuele Biggi / Harlequin frogs (Atelopus varius) in captivity, held in gloved hands at the El Valle de Anton Conservation Centre (EVACC), breeding program, Panama. Critically endangered.

Emanuele Biggi / Harlequin frogs (Atelopus varius) in vivarium in captivity at the El Valle de Anton Conservation Centre (EVACC), breeding program, Panama. Critically endangered.

Emanuele Biggi / Edgardo Griffith holding a Harlequin frog (Atelopus varius) in captivity at the El Valle de Anton Conservation Centre (EVACC), Panama. Critically endangered. EDITORIAL USE ONLY.

Emanuele Biggi / Edgardo Griffith holding a Harlequin Frog (Atelopus varius) in captivity at the El Valle de Anton Conservation Centre (EVACC), Panama. Critically endangered. EDITORIAL USE ONLY.

Emanuele Biggi / Harlequin frogs (Atelopus varius) females in captivity in vivaria at the El Valle de Anton Conservation Centre (EVACC), Panama. Critically endangered.

John Cancalosi / American toad (Anaxyrus americanus), two pairs mating. Maryland, USA. April.

David Perpinan / A pair of Natterjack toads (Epidalea calamita) mating in water, Cuenca, Spain, April. Filmed at night using an infrared camera.

David Perpinan / Male Natterjack toad (Epidalea calamita) calling, showing vocal sac inflating and then leaving frame, Cuenca, Spain, April. Filmed at night using an infrared camera.

David Perpinan / Natterjack toadlet (Epidalea calamita) resting, while other toadlets move around it, Barcelona, Spain, May.

David Perpinan / Rack focus between Natterjack toadlets (Epidalea calamita), Barcelona, Spain, May.

David Perpinan / Natterjack toadlet (Epidalea calamita) crawling over other toadlets, Barcelona, Spain, May.

David Perpinan / Natterjack toadlets (Epidalea calamita) moving across sand and stones, Barcelona, Spain, May.

David Perpinan / Natterjack toadlet (Epidalea calamita) investigating fly, Barcelona, Spain, May. - needs adusting

David Perpinan / Rack focus between a seed and Natterjack toadlet (Epidalea calamita), Barcelona, Spain, May.

David Perpinan / Natterjack toad (Epidalea calamita) gets kicked by mating rival, Barcelona, Spain, April.