Andy Rouse / Galapagos giant tortoise (Geochelone elephantopus / nigra) adult feeding, Santa Cruz, Galapagos

Tui De Roy / Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos.

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists, acting as ecosystem engineers by maintaining open meadows and digg...

Tui De Roy / Wolf giant tortoise (Chelonoidis becki) two males fight briefly over mating rights. Wolf Volcano, Isabela Island, Galapagos

Tui De Roy / Wolf giant tortoise (Chelonoidis becki) in habitat. Hybrids of mixed parentage with different shell shapes are scattered far and wide on the rugged west slope of Wolf Volcano. Wolf Volcano, Isabela Is...

Lucas Bustamante / Sierra Negra giant tortoise (Chelonoidis guntheri) Isabela Island, Galapagos Islands.

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) walking, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) pair mating in shallow pool, with two others resting nearby. Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoises (Chelonoidis porteri) resting in water, South coast (Las Palmitas), Santa Cruz Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoise (Chelonoidis porteri) feeding on grass, El Chato II, Highlands, Santa Cruz Island, Galapagos

Tui De Roy / Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos

Tui De Roy / Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species. Fausto Lle...

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) silhouetted at sunrise, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in pool, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in water at night, Alcedo Volcano, Isabela Island, Galapagos

Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) male mating with female while squabbling with another tortoise. Alcedo Volcano, Isabela Island, Galapagos

Andy Rouse / Alcedo volcano Galapagos giant tortoise (Chelonoidis vandenburghi) adult male on forest track, Urbina Bay, Isabela Island, Galapagos.

Tui De Roy / Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos

Tui De Roy / Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos

Tui De Roy / Hood island giant tortoise (Chelonoidis nigra hoodensis) used for breeding program since 1960s, Tortoise Breeding Centre, Puerto Ayora, Santa Cruz Island, Galapagos Islands, Ecuador, Critically endang...

Tui De Roy / Volcan Alcedo giant tortoises (Chelonoidis nigra vandenburghi) wallowing in rain pools on caldera floor, possibly for thermoregulation or detering parasites, with sulphur fumaroles in background, Alce...

Tui De Roy / Wolf-Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos. Did you know? Galapagos tortoises have a mutualistic relati...

Tui De Roy / Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait, Isabela Island, Galapagos

Pete Oxford / Saddleback form of Galapagos Giant Tortoise (Chelonoidis nigra) walking, Wolf Volcano, Isabela Island, Galapagos

Pete Oxford / Galapagos Giant Tortoise (Geochelone elephantophus / Chelonoidis nigra) head portrait, Charles Darwin Research Station. Puerto Ayora,Santa Cruz Island, Galapagos, endemic

Juan Carlos Munoz / Galapagos giant tortoise {Geochelone elephantopus} head portrait, Santa Cruz Island, Galápagos, January

Juan Carlos Munoz / Galapagos giant tortoise {Geochelone elephantopus} cooling in water, Santa Cruz Island, Galapagos, January

Pete Oxford / Galapagos Giant Tortoise (Geochelone elephantophus vandenburghi) near fumaroles, Alcedo Volcano crater floor, Isabela Island, Galapagos Islands

Tim Laman / Galapagos giant tortoise (Geochelone nigra / Geochelone elephantopus) feeding (captive). Charles Darwin Research Station, Santa Cruz Island, Galapagos Islands.

MORLEY READ / "Lonesome George", the last Galapagos Tortoise {Geochelone elephantopus} from Pinta Island, Galapagos

Pete Oxford / Galapagos Giant tortoise portrait {Geochelone elephantopus} Galapagos Is

Pete Oxford / D - Galapagos Giant tortoise portrait - Dome form {Geochelone elephantopus} retreating into shell. Darwin Research Station. Santa Cruz Is. Galapagos Islands, Ecuador. South America

Pete Oxford / Galapagos Giant tortoise close up portrait - dome form {Geochelone elephantopus} retreating into shell. Galapagos Is

Ingo Arndt / Giant tortoise {Geochelone elephantopus} Galapagos

Pete Oxford / Giant tortoise {Geochelone elephantopus}, Alcedo Volcano, Isabela Island, Galapagos

Pete Oxford / Giant Tortoise (Geochelone elephantopus) and tourists in the Galapagos Islands

Mark Moffett / Galapagos Giant Tortoise (Chelonoidis nigra) in Alcedo Volcano, Isabella Island, Galapagos Islands, Ecuador

Tui De Roy / Pinzon Island Tortoise (Chelonoidis nigra ephippium) pair fighting, Pinzon Island, Galapagos Islands, Ecuador

Tui De Roy / Saddleback Galapagos Tortoise (Chelonoidis nigra hoodensis) in defensive posture, Espanola Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) pair mating, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) on ridge, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) pair in volcanic landscape, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud at night, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador