Luke Massey / Spotted hyenas (Crocuta crocuta) in urban setting at night, Harar, Ethiopia

Neil Aldridge / Spotted hyaena (Crocuta crocuta) reassuring her tiny pup at the entrance to their den, Northern Tuli Game Reserve, Botswana.

Neil Aldridge / Spotted hyaena (Crocuta crocuta) exploring the skeleton of an elephant, Selinda Spillway, northern Botswana.

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) at night, taken with remote camera. Liuwa Plain National Park, Zambia. December

Will Burrard-Lucas / Spotted hyena (Crocuta crocuta) low angle shot of group taken with remote camera. Liuwa Plain National Park, Zambia. May

Enrique Lopez-Tapia / Man feeding Hyaenas (Crocuta crocuta) at night in Harar City, this has been a tradition for several centuries, and has now become a show for tourists. Ethiopia, November 2014

Enrique Lopez-Tapia / Man feeding Hyaenas (Crocuta crocuta) at night in Harar City, this has been a tradition for several centuries, and has now become a show for tourists. Ethiopia, November 2014

Solvin Zankl / Brown hyena (Hyaena brunnea) stalking fur seal (Arctocephalus pusillus) pup, Sperrgebiet National Park, Namibia, December. Sequence 1/8

Anup Shah / Spotted hyena (Crocuta crocuta) walking and smelling the ground. Maasai Mara National Reserve, Kenya.

Anup Shah / White-backed vultures (Gyps africanus) waiting for Spotted hyenas (Crocuta crocuta) to finish feeding on a kill. Maasai Mara National Reserve, Kenya.

Anup Shah / Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) aged about 2 years watching with curiosity with the rest of the clan in the background. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) pup approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyenas (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Spotted hyena (Crocuta crocuta) approaching with curiosity. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Denis-Huot / Spotted hyena (Crocuta crocuta) hunting Lesser flamingoes (Phoenicopterus minor) Lake Nakuru, Kenya

Denis-Huot / Spotted hyena (Crocuta crocuta) baby at den, Masai-Mara Game Reserve, Kenya

Martin Dohrn / Three hyenas (Crocuta crocuta) approaching a resting lion (Panthera leo) at night. Topi Plain, Masai Mara, Kenya. Image taken using an infared camera with no artificial visible light, on location for...

Anup Shah / Spotted hyena (Crocuta crocuta) portrait standing, Masai Mara National Reserve, Kenya. December

Anup Shah / Spotted hyena (Crocuta crocuta) pups approaching with curiosity, Masai Mara National Reserve, Kenya. December

Anup Shah / Spotted hyena (Crocuta crocuta) approaching with suspicion, Masai Mara National Reserve, Kenya. December

Laurent Geslin / Brown Hyaena (Hyaena brunna) running. Skeleton Coast, Namibia

Anup Shah / Female Spotted hyenas (Crocuta crocuta) walking through savanna, Masai Mara National Reserve, Kenya. March

Anup Shah / Spotted hyena pups greeting an adult (Crocuta crocuta). Masai Mara National Reserve, Kenya. December

Anup Shah / Spotted hyaena {Crocuta crocuta} chasing Lesser flamingo {Phoeniconaias minor} Lake Nakuru NP, Kenya

Anup Shah / Spotted hyaena {Crocuta crocuta} amongst flock of Lesser flamingo {Phoeniconaias minor} Lake Nakuru NP, Kenya

Anup Shah / Spotted hyaena {Crocuta crocuta} amongst flock of Lesser flamingo {Phoeniconaias minor} Lake Nakuru NP, Kenya

Christophe Courteau / Spotted Hyaena (Crocuta crocuta) at night, South Africa

Anup Shah / Spotted hyaena {Crocuta crocuta} running with prey of Thomson's gazelle fawn, Serengeti NP, Tanzania

Anup Shah / Spotted hyaena {Crocuta crocuta} at dusk walking to Acacia tree. Maasai Mara, Kenya.

Anup Shah / Spotted hyaena {Crocuta crocuta} chasing vulture from carcass. Maasai Mara, Kenya.

Anup Shah / Spotted hyaena chases another with gazelle prey, Serengeti NP, Tanzania {Crocuta crocuta}

Anup Shah / Spotted hyaena predating newborn Thomson's gazelle, Serengeti NP, Tanzania {Crocuta crocuta}

Richard Du Toit / Spotted hyaena {Crocuta crocuta} with Buffalo bone. Moremi WR Botswana

Richard Du Toit / Spotted hyaenas greeting at dusk {Crocuta crocuta} Okavango delta, Botswana

Martin Dohrn / Spotted hyaenas {Crocuta crocuta} on the prowl. Serengeti NP, Tanzania, East Africa. Image taken using 'Starlight Camera' technology without artificial lighting. Ngorongoro Conservation Area,

Martin Dohrn / Spotted hyaenas harrass lioness. Serengeti NP, Tanzania, East Africa. Infra-red image taken at night using 'Starlight Camera' technology without artificial lighting. Ngorongoro Conservation Area,

Martin Dohrn / Spotted hyaena portrait in moonlight {Crocuta crocuta} Serengeti NP Tanzania, East Africa. Image taken using 'Starlight Camera' technology without artificial lighting. Ngorongoro Conservation Area,

Martin Dohrn / Spotted hyaenas {Crocuta crocuta} feeding on kill, Serengeti NP, Tanzania, East Africa. Image taken at night using 'Starlight Camera' technology without artificial lighting. Ngorongoro Conservation Ar...

Martin Dohrn / Hyaenas threatening lions who have stolen kill. Serengeti NP, Tanzania, East Africa. Image taken using 'Starlight Camera' technology and infra red light. Ngorongoro Conservation Area

Anup Shah / Spotted hyaena {Crocuta crocuta} carries Wildebeest head Masai Mara GR, Kenya

Anup Shah / Spotted hyaena {Crocuta crocuta} greeting ritual, Masai Mara, Kenya East-Africa

Anup Shah / Spotted hyaena greeting ritual {Crocuta crocuta} Masai Mara, Kenya East-Africa

Martin Dohrn / Spotted hyaenas at night, Masai Mara, Kenya. Starlight image intensifier camera image taken with no artificial light, composite

Terry Andrewartha / Spotted hyaenas {Crocuta crocuta} taunting lion Maasai Mara Game Reserve, Kenya, East Africa

Karl Ammann / Male Lion killing Hyaena, Masai Mara, Kenya

Constantinos Petrinos / Spotted hyena cubs {Crocuta crocuta} Masai Mara, Kenya, Africa

Anup Shah / Spotted hyaena and White backed vulture dispute at carcass, Masai Mara, Kenya