Will Burrard-Lucas / A mating pair of leopards (Panthera pardus) court in front of a camera trap at night, Laikipia Wilderness Camp, Kenya. Photographed with a camera trap. EDITORIAL USE ONLY. All other uses require clear...
Simon Littlejohn / Juvenile Lammergeier vulture (Gypaetus barbatus) in flight, mobbed by a raven, Grand Bargy cirque, Haute Savoie, France, October.
Simon Littlejohn / Bonelli's eagle chicks (Aquila fasciata) in nest pecking each other, aged one week, Andalusia , Spain, May.
Tui De Roy / Black rhino (Diceros bicornis) and White rhino (Ceratotherium simum), being harrassed by playfighting White rhino bull, Solio Game Reserve, Laikipia, Kenya. September.
Matthew Maran / Red fox (Vulpes vulpes) family in an allotment, with rival male fox in background, London, England, UK, June.
Matthew Maran / Male Red fox (Vulpes vulpes) interacting with cub in allotment, London, England, UK, June.
David Hall / Steller sea lions (Eumetopias jubatus), three playing underwater, Inian Island, Alaska, USA. August.
David Hall / Steller sea lions (Eumetopias jubatus) playing underwater, Inian Island, Alaska, USA. August.
John Cancalosi / Grey seal (Halichoerus grypus), mother and young interacting on beach, UK
Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. June.
Matthew Maran / Red fox (Vulpes vulpes) dog interacting with a vixen in an urban garden. North London, UK. July.
Morley Read / Slow motion clip of a group of male Mallards (Anas platyrhynchos) competing violently to mate with a single female, Cornwall, England, UK, May 2018.
Nick Upton / Marbled white butterflies (Melanargia galathea) nectaring on a clump of Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, England, UK, June.
Ben Gillett / Slow motion clip of a female Appaloosa horse grooming a male, Bristol, England, UK, May.
John Chan / Two male Southern Pacific rattlesnakes (Crotalus oreganus helleri) fighting during mating season. This behaviour is known as a "combat dance", and is used to determine dominance and the right to mate...
Terry Whittaker / RF - Arctic foxes (Vulpes lagopus) interacting Blue colour morph. Hornstrandir Nature Reserve, Iceland. March. (This image may be licensed either as rights managed or royalty free.)
Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
Marie Read / Western Grebe (Aechmorphus occidentalis), pair during 'rushing' courtship display in which they run across water's surface in synchrony, near Escondido, California, USA, January.
Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.
Espen Bergersen / Aerial shot of a female Humpback whale (Megaptera novaeangliae) and calf surrounded by Killer whales (Orcinus orca), mother hits sideways with her tail fluke to drive Killer whales away, Skjervoy, Tro...
David Gallan / Two Southern right whales (Eubalaena australis) playing in shallows, taken with drone, Tathra Beach, New South Wales, Australia, August.
Loic Poidevin / Northern ravens (Corvus corax) interacting in snow, Leon, Spain, February.
John Chan / Western gulls (Larus occidentalis) attacking an Elegant tern (Thalasseus elegans) with a broken wing, Southern California, USA, April.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
David Perpinan / Group of Griffon vultures (Gyps fulvus) fighting to feed on a dead lamb. Cuenca, Spain, August.
David Perpinan / Group of Griffon vultures (Gyps fulvus) arriving and feeding on a dead lamb, Cuenca, Spain, August.
David Perpinan / Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, Barcelona, Spain, May.
David Perpinan / Grey heron (Ardea cinerea) chick trying to force its parent to regurgitate food, with another chick leaning in, chased away by adult, Barcelona, Spain, May.
Terry Whittaker / Juvenile Red fox (Vulpes vulpes) interacting with and trying to play with an adult in an urban garden, Greater Manchester, UK, August. Filmed using a camera trap.
Terry Whittaker / Domestic cat chasing a Red fox (Vulpes vulpes) in an urban garden, Greater Manchester, UK, August 2017. Filmed using a camera trap.
Dave Bevan / Slow motion clip of two pairs of Jackdaws (Corvus monedula) squabbling over nest site, Worcestershire, England, UK, May.
Five Films / Female Grey seal (Halichoerus grypus) defending pup on beach, chasing male away, Horsey, Norfolk, November.
David Perpinan / Two Hyacinth macaws (Anodorynchus hyacinthus) in a tree, Pantanal, Mato Grosso do Sul, Brazil.
John Waters / Pair of Brown trout (Salmo trutta) spawning, with others eating eggs, River Kennet, Berkshire, England, UK. March.
John Waters / Female Grayling (Thymallus thymallus) pursued by several males, River Kennet, Berkshire, England, UK. March.
John Waters / Pair of Brown trout (Salmo trutta) spawning, with another male trying to fertilise eggs River Kennet, Berkshire, England, UK. February.
Francisco Marquez / Griffon vultures (Gyps fulvus) and European black vultures (Aegypius monachus) feeding on a dead deer, Cabaneros National Park, Montes de Toledo, Spain, May.
SCOTLAND: The Big Picture / Red squirrels (Sciurus vulgaris) interacting, Cairngorms National Park, Highlands, Scotland, UK, January.
Brian Bevan / Brown hare (Lepus europaeus) boxing, Bedfordshire, England, UK, March.
Jabruson Motion / Male Western gorilla (Gorilla gorilla) displaying, throws clump of grass into the air, runs through bai and hits other gorilla, Maya Nord Bai, Odzala-Kokoua National Park, Cuvette-Ouest Region, Republ...
Jabruson Motion / Close-up of a male Mountain gorilla (Gorilla beringei beringei) looking at camera, pulls off lens component, Bukima, Virunga National Park, Democratic Republic of Congo, 1996.
Dave Bevan / Two male Grey seals (Halichoerus grypus) fighting in surf, with a female and juvenile nearby, Norfolk, England, UK. December.
Francisco Marquez / Pair of Iberian lynxes (Lynx pardinus) greeting, Andalusia, Spain, January.
Christophe Courteau / Female Mountain gorilla (Gorilla beringei beringei) playing with baby, Volcanoes National Park, Rwanda.
Sandesh Kadur / Female Dussumier's malabar langur (Semnopithecus dussumieri) feeding with baby, picks it up and walks away, Ranthambore National Park, Rajasthan, India. 2016.
Sandesh Kadur / Two Bengal monitors (Varanus bengalensis) fighting, Ranthambore National Park, Rajasthan, India. 2016.
David Tipling / King penguins (Aptenodytes patagonicus) pair interacting, part of courtship, Gold Harbour South Georgia, January