Nick Upton / Great capricorn / Great oak longhorn beetle (Cerambyx cerdo), one of the largest beetles in Europe, on tree trunk, Lesbos (Lesvos), Greece, June.

Nick Upton / Shoal of young sardines (Sardina pilchardus) swimming in various directions close to the shore in a shallow sandy bay, viewed from above water. Eastern Lesbos /Lesvos, Greece, August.

Nick Upton / Shoaling young sardines (Sardina pilchardus) swimming in shallow water in a sandy bay, casting shadows on the seafloor, viewed from above water. Eastern Lesbos / Lesvos, Greece, August.

George McCarthy / Bittern (Botaurus stellaris) standing in grass. Lesvos, Greece, Europe

Ashley Cooper / Life jackets and remains of boats left by Syrian migrants, refugees fleeing the war. Lesvos Island, Efthalou, Greece. September.

Ashley Cooper / Syrian migrants, refugees fleeing the war, Lesvos Island, Greece. September 2015.

Ashley Cooper / Life jackets and remains of boats left by Syrian migrants, refugees fleeing the war. Lesvos Island, Efthalou, Greece. September.

Ashley Cooper / Solar thermal panels for heating water in Skala Eresou on Lesvos, Greece. June 2013

Ashley Cooper / A swimming pool at a holiday complex in Skala Eresou on Lesvos, Greece. June 2013

Ashley Cooper / Striped necked terrapins (Mauremys caspica) in a river in Skala Eresou, Lesvos, Greece. June.

Ashley Cooper / Beech marten, (Martes foina) killed on a road on Lesvos, Greece. June.

Ashley Cooper / Beech marten, (Martes foina) killed on a road on Lesvos, Greece. June.

Ashley Cooper / An old windmill converted into a house in Sigri on Lesvos, with a fossilised tree in the foreground. Greece, June 2013

Ashley Cooper / Wind farm above Eressos on Lesvos, Greece. June 2013

Nick Upton / White-spotted rose beetles (Oxythyrea funesta) mating on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Turkish meadow brown butterfly (Maniola megala) feeding on Cretan oregano (Origanum onites) flowers with wings closed, Lesbos (Lesvos), Greece, May.

Nick Upton / Tumbling flower beetle / Pintail beetle (Mordellidae) feeding on Cretan oregano flower (Origanum onites) Lesbos / Lesvos, Greece, May.

Nick Upton / Large thistle weevil / Sluggish weevil (Cleonis pigra), on fabric, Lesbos / Lesvos, Greece, May.

Nick Upton / Thick-headed fly (Physocephala rufipes), a parasite of bees, feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Sweat bee (Halictus cf. brunnescens) female foraging on Cretan organo (Origanum onites), Lesbos/ Lesvos, Greece.

Nick Upton / Spurge hawkmoth caterpillar (Hyles euphorbiae) on Broad-leaved glaucous spurge / Myrtle spurge (Euphorbia myrsinites) stripped of its leaves, on limestone mountain top, Mount Olympus, Lesbos / Lesvos,...

Nick Upton / Spurge hawkmoth caterpillar (Hyles euphorbiae) feeding on Sea spurge (Euphorbia paralias) among sand dunes, Lesbos / Lesvos, Greece, May.

Nick Upton / Thread-winged lacewing / Spoonwing lacewing (Nemoptera sinuata) feeding on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Female Greek paper wasps (Polistes hellenicus) tending grubs in their nest in coastal scrubland, Lesbos / Lesvos, Greece, May.

Nick Upton / Femae Greek paper wasps (Polistes hellenicus) on their nest in coastal scrubland, Lesbos / Lesvos, Greece, May.

Nick Upton / Female Napoleon crab spider (Synema globosum) waiting for prey on Cretan oregano flowerhead (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mediterranean flatheaded woodborer / Jewel beetle (Capnodis cariosa / tenebrionis), a pest of fruit trees including Apricot and Almond, well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May

Nick Upton / Mammoth wasp / Giant solitary wasp (Megascolia maculata maculata) female sunning on a tree trunk, Lesbos/ Lesvos, Greece, May.

Nick Upton / Lynx spider (Oxyopes heteropthalmus) hunting for insect prey on Cretan oregano flowers (Origanum onites), Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a thistle, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flower, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Long-legged bush cricket nymph (Acrometopa syriaca or servillea) standing on a Broad-leaved Glaucous Spurge / Myrtle spurge (Euphorbia myrsinites) flower, Mount Olympus, Lesbos / Lesvos, Greece, May.

Nick Upton / Male Ladybird spider (Eresus cinnaberinus / niger) searching for females, Lesbos / Lesvos, Greece, May.

Nick Upton / Male Ladybird spider (Eresus cinnaberinus / niger) searching for females on a rock face, Lesbos / Lesvos, Greece, May.

Nick Upton / Horse fly (Pangonius funebris), a carpenter bee mimic, using its long proboscis to forage on Milk thistle (Carduus marianus), Lesbos / Lesvos, Greece, May.

Nick Upton / Hornet hoverfly / Banded hoverfly (Volucella zonaria) feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Hornet hoverfly / Banded hoverfly (Volucella zonaria) feeding on Japanese mock orange flowers (Pittosporum tobira), Lesbos/ Lesvos, Greece, May.

Nick Upton / Heather crab spider (Thomisus onustus) waiting for insect prey on Cretan oregano flowerhead (Origanum onites), Lesbos/ Lesvos, Greece, May.

Nick Upton / Heather crab spider (Thomisus onustus) waiting for insect prey on Cretan oregano flowerhead (Origanum onites), Lesbos/ Lesvos, Greece, May.

Nick Upton / Female Harvestman (Zacheus crista) well camouflaged on a tree trunk, Lesbos / Lesvos, Greece, May.

Nick Upton / Female Harvestman (Zacheus crista) on a house wall, Lesbos / Lesvos, Greece, May.

Nick Upton / Digger wasp (Sphex pruinosus) male sunning on a rock in hot sunshine and raising its legs, Lesbos/ Lesvos, Greece, May

Nick Upton / Digger wasp (Sphex pruinosus) male sunning on a rock in hot sunshine and raising its legs, Lesbos/ Lesvos, Greece, May

Nick Upton / Copper Buprestid / Jewel beetle (Aurigena / Perotis lugubris) a pest of fruit tee roots, Lesbos / Lesvos, Greece, May

Nick Upton / Copper Buprestid / Jewel beetle (Aurigena / Perotis lugubris) a pest of fruit tee roots, Lesbos / Lesvos, Greece, May

Nick Upton / Young Western Caspian pond turtle / Balkan stripe-necked terrapin (Mauremys caspica rivulata) swimming in a pond with its head above water, Isle of Lesbos / Lesvos, Greece, May.