Ross Hoddinott / Westward Ho! beach at sunrise, colourful sky at low tide and tidal pool, north Devon, UK. February.
Theo Bosboom / Star shaped sand pattern as photographed at low tide at Luskentyre beach, Lewis and Harris, Scotland, UK, October.
John Chan / Western gull (Larus occidentalis) feeding on a California two-spot octopus (Octopus bimaculoides) on a mudflat at low tide, Southern California, USA, November.
Duncan Murrell / Coral formations at low tide in Prony Bay in the Southern Lagoon, Lagoons of New Caledonia: Reef Diversity and Associated Ecosystems UNESCO World Heritage Site. New Caledonia.
Jurgen Freund / Timelapse of Periwinkle snails (Littoraria) climbing up Mangrove (Rhizophora) roots at lowtide, Daintree, Wet Tropics World Heritage Area, North Queensland, Australia.
Pedro Narra / Aerial view of low tide in Sado estuary, Portugal. November
Alex Hyde / Colony of Common Mussels (Mytilus edulis) growing on striated rock formation exposed at low tide. Cornwall, England, UK. Highly commended in the Coast and Marine Category of the British Wildlife Photo...
Theo Bosboom / Limpets (Patellidae) exposed at low tide on the coast of Playa de Ursa, Portugal. Finalist in the Animals in their Environment Category of the Wildlife Photographer of the Year Awards (WPOY) Competiti...
Guy Edwardes / View over Pednvounder Beach towards Logan Rock, Porthcurno, Cornwall, England, UK. April 2009.
Ernie Janes / Low tide on the Wash, Snettisham, Norfolk, England, UK, February.
Steven Kazlowski / Polar bear (Ursus maritimus) curious young bear approaching camera, over newly forming pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Ernie Janes / Oystercatcher (Haematopus ostralegus) footprints on sand at low tide, Norfolk, England, UK, November.
Peter Cairns / Isle of Rum at sunset viewed from Eigg, Scotland, April 2013.
Charlie Summers / Black Bear (Ursus americanus) hunting for spawning Salmon during low tide, Neets Creek estuary, Alaska, July.
Tom Gilks / Zanzibari woman at low tide with harvested seaweed (eucheuma spinosum) Matemwe, Zanzibar, Tanzania
Ernie Janes,Ernie Janes / RSPB Nature reserve at Titchwell Marsh Norfolk, May
Merryn Thomas / Man taking a photograph of Three Cliffs Bay, Gower, Swansea, Wales. January 2013. Model released.
Graham Eaton / Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea at sunrise, Cayenne, French Guiana, July
Ernie Janes / Holkham beach at low tide, Norfolk, UK August
Nick Upton / Common Shore crab (Carcinus maenas) on clump of Toothed wrack (Fucus serratus) exposed on shore at low tide, St. Bees, Cumbria, UK, July.
Alex Mustard / 2020VISION / Looking up through water at boy dipping in a rock pool with a shrimping net. Falmouth, Cornwall, England, UK. Model released, July
Alex Mustard / 2020VISION / Children enjoying dipping in rockpools at low tide in Falmouth, Cornwall, England, UK, July, all model released
Ingo Arndt / King Penguin (Aptenodytes patagonicus) dispute on beach, Volunteer Point, Falkland Islands, February
Merryn Thomas / View across Oxwich Bay on the Gower Peninsula, South Glamorgan, Wales, January 2012.
Peter Cairns / 2020VISION / Old fishing boat held upright at low tide with ropes and chains, Roa Island, Morecambe Bay, Cumbria, England, UK, February 2012. Did you know? Over the last decade the number of UK fishermen has decl...
Philippe Clement / Medieval fish lock at low tide on the island Ile de Re, Charente-Maritime, France September 2011
Alex Mustard / 2020VISION / View up through tidepool of 2020Vision young photographer, Bertie Gregory, Falmouth, Cornwall, UK, July 2011. Model released. Did you know? Life isn’t easy in a rock pool! Animals here must deal with...
Ernie Janes / Gulls on the beach at sunset, The Wash, Hunstanton, Norfolk, UK, September 2011
Andrew Parkinson / Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October.
Fiona Rogers / Proboscis Monkey (Nasalis larvatus) male walking on the mudflats of a mangrove swamp revealed at low tide. Bako National Park, Sarawak, Borneo, Malaysia, March.
Anup Shah / Proboscis monkey (Nasalis larvatus) male walking across the mudflats of a mangrove swamp revealed at low tide with the sea in the background, Bako National Park, Sarawak, Borneo, Malaysia, March
Terry Whittaker / 2020VISION / Common / Harbour seals (Phoca vitulina) hauled out on mudflats at saltmarsh, Wallasea Island RSPB reserve, Essex, UK, March 2011
Peter Cairns / 2020VISION / Two women walking along sandy beach at low tide, near Maryport, Solway Firth, Cumbria,UK, April 2011
Matthew Maran / Surfer Benj Yougson at Long Beach. Pacific Rim National Park Reserve, west coast of Vancouver Island, Canada, September 2010. Model released
Steve Nicholls / Horseshoe crabs (Limulus polyphemus) spawning at low tide, Slaughter Beach, Delaware, USA, May
Inaki Relanzon / Woman and girl fishing on the beach. Morondava, Madagascar, Africa.
Eric Baccega / Flowerpot sea stacks with outgoing tide at Hopewell Rocks. The world's highest tides. Bay of Fundy, New Brunswick, Canada, September 2010
Ernie Janes / English Springer spaniel running over beach at low tide, August, UK
Edwin Giesbers / Two Long-tailed / Crab-eating macaques (Macaca fascicularis) foraging on coastline at low tide, Bako National Park, Sarawak, Borneo, Malaysia
Brandon Cole / Woman (model released) explores tidepools exposed during a summer's extreme low tide. Split level view of a tidepool showing Ochre sea stars (Pisaster ochraceus), Giant green sea anemones (Anthopleura...
Brandon Cole / Woman (model released) explores tidepools exposed during a summer's extreme low tide. Split level view of a tidepool, marine life includes Ochre sea stars (Pisaster ochraceus), Purple sea urchins (Str...
David Woodfall / Severn Estuary at ebb tide in rainstorm. This habitat is potentially threatened by proposed tidal barrage. England, UK April 2010
Anup Shah / Troop of Proboscis Monkeys (Nasalis larvatus) walking on their hindlegs through the mudflats revealed at low tide. Bako National Park, Sarawak, Borneo, Malaysia, March 2010.
Roger Powell / Bar-tailed godwit (Limosa lapponica) in winter plumage, extracting a Lugworm from a deep hole at low tide, Boulmer, Northumberland, UK
Ross Hoddinott / Porthcothan Bay, evening light and sunset at low tide. Near St Eval, Cornwall, England, UK. April 2010.
Wild Wonders of Europe / LÃ³pez / Aerial view of saltmarsh exposed at low tide, Bahía de Cádiz Natural Park, Cádiz, Andalusia, Spain, March 2008
Adam Burton / Bedruthan Steps, at low tide with stormy sky, Cornwall, England. May 2009
Ross Hoddinott / Burgh Island and Burgh Island hotel. Evening light at low tide. Taken from Bigbury Bay (Blue Flag beach) South Devon, UK. September 2009.