Andres M. Dominguez / Holm oak (Quercus ilex) in the Sierra de Grazalema Natural Park, Benaocaz, southern Spain, November.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) colony huddled together under the Southern Lights, Aurora australis, Atka Bay, Antarctica. September.
Ross Hoddinott / Marbled white butterfly (Melanargia galathea) resting among tall grasses and bathed in warm, early morning sunlight, Volehouse Moor, Devon, UK. July.
Ross Hoddinott / Marbled white butterfly (Melanargia galathea) roosting among tall grasses at sunrise and bathed in warm, early morning sunlight, Volehouse Moor, Devon, UK. July.
Ernie Janes / Beech trees (Fagus sylvatica) at night in winter, with lamp post illuminating the woodland. Felbrigg Woods, Norfolk, England, UK. January.
Claudio Contreras / Puente de Dios cascades, Tamasopo, Huasteca Region, Mexico, August
Guy Edwardes / Sgwd Ddwli Uchaf, Brecon Beacons National Park, Wales. June 2013
Jack Dykinga / Aurora Borealis reflected in Polar Lake, conifers silhouetted at edge. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Jack Dykinga / Aurora Borealis display above conifers and Polar Lake. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Jack Dykinga / Aurora Borealis reflected in Polar Lake, wetland and conifers silhouetted along shore. Near Great Slave Lake, Northwest Territories, Canada. September 2018.
Bernard Castelein / Waterviolet / featherfoil (Hottonia palustris) at sunset, Groot Schietveld, Wuustwezel, Belgium
Bernard Castelein / Sunrise reflected in water with ripples, Klein Schietveld, Brasschaat, Belgium
Bernard Castelein / Scots pine tree (Pinus sylvestris) at sunrise, Klein Schietveld, Brasschaat, Belgium. September.
Bernard Castelein / Frozen water patterns in Klein Schietveld, Brasschaat, Belgium
Bernard Castelein / Sunrise over Groot Schietveld, Wuustwezel, Belgium. June 2015.
Bernard Castelein / Scots pine tree (Pinus sylvestris) with sunrays at sunrise. Klein Schietveld, Brasschaat, Belgium.
Theo Bosboom / Giants Causeway at night, lit by full moon. County Antrim, Northern Ireland. Winner of the Landscapes Category of the GDT European Wildlife Photographer of the Year 2017.
SCOTLAND: The Big Picture / Scots pine (Pinus sylvestris) with Northern lights / Aurora borealis lighting up the night sky in background, Monadhliath Mountains, Cairngorms National Park, Scotland, UK, October 2015.
Andy Rouse / Great crested grebe (Podiceps cristatus) swimming, backlit with morning light forming a bokeh affect on the water, Wales, UK, February. Highly commended in the Habitat Category of the BWPA (British W...
Bence Mate / Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May.
Georgette Douwma / Upside down jellyfish (Cassiopeia andromeda) Lembeh, Sulawesi, Indonesia.
Erlend Haarberg / Mountain hare (Lepus timidus) backlit in snow, Vauldalen, Norway, May.
Peter Lewis / Super moon rising behind Glastonbury Tor with people watching, Somerset, England, UK, 27th September 2015.
Wim van den Heever / Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June.
Wim van den Heever / Quiver tree forest (Aloe dichotoma) at night with stars and the Milky Way, Keetmanshoop, Namibia.
Alex Mustard / Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Sven Zacek / Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.
Sven Zacek / Aerial view at sunrise of fog over bog forest, Tartumaa, Estonia, May 2013.
SCOTLAND: The Big Picture / Pod of Bottlenose dolphins (Tursiops truncatus) surfacing just offshore, Black Isle, Scotland, UK, July.
Andy Rouse / Sunrise in forest, silhouetting trees. Ranthambhore Tiger Reserve, India.
Ross Hoddinott / Corfe Castle, early morning llight and mist, Corfe Castle, Dorset, UK. September 2014.
Peter Cairns / Northern lights (Aurora borealis) over Skagsanden beach, Flakstadoya, Lofoten, Norway. March 2015.
Peter Cairns / Svalbard reindeer (Rangifer tarandus platyrhynchus) backlit in evening light, Spitsbergen, Svalbard, Norway, August.
Mark MacEwen / Firefly squid (Watasenia scintillans) close to beach during mating season. Toyama Bay, Japan.
Mark MacEwen / Japanese firefly (Luciola cruciata) light trails at night above stream in rural Japan. June.
Yukihiro Fukuda / Light trails of Japanese fireflies (Hotaria parvula and Luciola cruciata), Nichinan-chou, Tottori, Japan, July, Endemic species.
Jurgen Freund / Wangi Falls at night illuminated by moonlight, Litchfield National Park, Northern Territory, Australia. December 2012.
Ole Jorgen Liodden / Barn swallow (Hirundo rustica) in mud collecting nesting material, collecting nesting material, with golden bokeh affect in puddle, Svalbard, Norway, June.
Ole Jorgen Liodden / Chatham stream on the main Chatham island, New Zealand, March 2014.
Russell Cooper / Blackening waxcap fungus (Hygrocybe sp.) New Forest, Hampshire, UK.
Andy Rouse / Red deer (Cervus elaphus) stag silhouetted at dusk, Wales, UK. Captive in deer park.
Doug Perrine / Rough-toothed dolphins (Steno bredanensis) Keauhou, Kona, Hawaii, USA, Central Pacific Ocean.
Floris van Breugel / Aurora Borealis over yellow tent lit from within and mountainous scenery, Arrigetch Peaks, Gates of the Arctic National Park, Brooks Range, Alaska, USA, August 2014.
Floris van Breugel / Ice cave melting in Mendenhall Glacier, Juneau, Alaska, USA, August 2014.
Floris van Breugel / Ice cave with melt water running through, Mendenhall Glacier, Juneau, Alaska, USA, August 2014.
Pal Hermansen / Aerial view of river delta, South West Iceland, June 2014.
David Pattyn / Great crested grebe (Podiceps cristatus) adult bird backlit with bokeh affect in foreground, The Netherlands.May 2014
David Pattyn / Coot (Fulica atra) adult close-up. Backlit at dawn. The Netherlands. April 2014