Martin Dohrn / Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using 'Starlight Camera' technology without artificial lighting.
John Shaw / King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November.
Alex Hyde / Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November.
Daniel Heuclin / Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018.
TJ Rich / Four chocolate working cocker spaniels on wall. Mother and offspring. Monmouth, Monmouthshire, Wales
TJ Rich / Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
Tui De Roy / Blue-footed booby (Sula nebouxii), five diving into sea. Espumilla Beach, Santiago Island, Galapagos.
David Tipling / Eurasian magpies (Pica pica) gathering prior to going to roost Hortobagy National Park, Hungary January
Dave Bevan / Slow motion clip of a Robin (Erithacus rubecula) flying to nest and feeding chicks, removes faecal sac, Carmarthenshire, Wales, UK, May.
David Perpinan / Mallards (Anas platyrhynchos) paddling in river, seen from underwater, Scotland, UK, April.
Edwin Giesbers / RF - Banded darter (Sympetrum pedemontanum) group of five resting, Netherlands, September. (This image may be licensed either as rights managed or royalty free.)
Yashpal Rathore / Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December.
Wim van den Heever / Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Laurie Campbell / Common / Viviparous lizard (Zootoca vivipara) mother with young. Burnmouth, Berwickshire, England, UK.
Tony Wu / Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March.
Doug Perrine / False killer whale (Pseudorca crassidens) pod, off the Kona Coast of Hawaii Island, Hawaiian Islands, USA. January.
Bence Mate / Great egret (Ardea alba) group of five at water's edge, with very out of focus egret in foreground. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary, February.
Bence Mate / Group of five Brown-hooded parrots (Pyrilia haematotis) in rain, Costa Rica.
Wim van den Heever / Blue wildebeest (Connochaetes taurinus) herd silhouetted against the rising sun with clouds in the background. Greater Kruger National Park, South Africa
Wim van den Heever / Plains zebra (Equus quagga) group of four including foal at sunset, Savuti Marsh, Botswana.
Jeff Vanuga / White horses of the Camargue galloping through marshes in the Camargue, France. April.
Jordi Chias / Bryde's whale (Balaenoptera edeni) and Common dolphins (Delphinus delphis) chasing small fish offshore. Santa Maria Island, Azores, Atlantic Ocean. Third Place in the Underwater Category of the Por El...
Jabruson / Mozambican children with captive Yellow Baboon youngster (Papio cynocephalus) young animal caught during troop crop raiding. Pemba to Montepuez highway, north-eastern Mozambique, November 2011. Specia...
Pal Hermansen / Inuit sled dogs resting, near Ittoqortoormiit, Scoresbysund, Greenland. Second Prize in the Nature category of the Spider Black and White Photography Competition 2015.
Staffan Widstrand / Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa. Specially commended in the Animal Portraits category of...
Klein & Hubert / Meerkat (Suricata surricatta) group of five warming up in the sunshine, Kalahari desert, South Africa.
Alex Mustard / Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey, and is followed by a Napolean wrasse (Cheilinus undulatus) and a Bluef...
Bence Mate / Great egrets (Ardea alba) and Grey heron (Ardea cinerea) squabbling over fish, with some in flight. Blurred motion photograph, Lake Csaj, Pusztaszer, Hungary. February.
Pascal Pittorino / Corsican pig (Sus scrofa domestica) sow, suckling the piglets, Parc Naturel Regional de Corse / Natural Regional Park of Corsica, Corsica Island, France. September.
Jabruson Motion / Group of Chimpanzees (Pan troglodytes) playing in an enclosure and drinking from bottles, Lwiro Primate Rehabilitation Center, South Kivu, Democratic Republic of Congo. Captive.
Loic Poidevin / Common teasel (Dipascus fullonum) seed-heads covered in ice, Vendee, France , January
Cyril Ruoso / Ringed-tailed lemur (Lemur catta) foraging in sisal field looking for flowers during a poor year without much fruit. Berenty reserve, Madagascar, September.
Cyril Ruoso / Keeper feeding Common woolly monkeys (Lagothrix lagotricha) in Ikamaperou Sanctuary, Amazon, Peru. October 2006.
Cyril Ruoso / Vicuna (Vicugna vicugna) herd in Altiplano, Aguada Blanca National Reserve, Peru.
Cyril Ruoso / African elephant (Loxodonta africana) hanging upside down, during translocation by Kenya Wildlife Service, due to over population, Mwaluganje Reserve, Kenya. October 2006.
Sven Zacek / Four Roe deer (Capreolus capreolus) bucks with velvet antlers and the shadow of a fifth buck, Tartumaa, Estonia, March.
Roger Powell / Small flock of Brent Geese (Branta bernicla) in flight..Texel Island, The Netherlands, Europe
Staffan Widstrand / Kittiwakes (Rissa tridactyla) nesting on wall of house, Vardo town, Varanger Peninsula, Norway, March.
Andy Rouse / Magnificent frigate bird (Fregata magnificens) group flying against cloudy sky, Galapagos.
George Sanker / American buffalo (Bison bison) with group of calves, Yellowstone National Park, Wyoming, USA. May.
Juan Manuel Borrero / Argan tree (Argania spinosa) fruits, cultivated plant, endemic to Morocco.
Juan Manuel Borrero / Wind turbines, Collet dels Feixos, Tarragona, Catalonia, Spain, May 2013.
Staffan Widstrand / Wild Wonders of China / Crested myna starlings (Arcidotheres cristatellus) sitting on domestic waterbuffaloes (Bubalus domesticus), Poyang Ho Lake, Jiangxi province, China
Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July
Chris & Monique Fallows / Long-beaked common dolphin (Delphinus capensis) school near the water?s surface, False Bay, South Africa
Enrique Lopez-Tapia / Hamer women in traditional clothing at the Jumping of the bulls ceremony. Ethiopia, November 2014
Denis-Huot / African elephant (Loxodonta africana) herd eating, Masai-Mara Game Reserve, Kenya
Pal Hermansen / Graveyard with white crosses half buried in snow, Svalbard, Norway. March.