Silvia Reiche / Brown Hairstreak (Thecla betulae) butterfly resting on blade of grass, Netherlands
Duncan Usher / Eurasian Spoonbill (Platalea leucorodia), Alentejo, Portugal
Silvia Reiche / Oldworld Swallowtail (Papilio machaon) butterfly on red autumn leaf, Hoogeloon, Noord-Brabant, Netherlands
Jan Wegener / Common Nighthawk (Chordeiles minor) male in display flight, Winnie, Texas
Staffan Widstrand / Wild Wonders of China / Banyan fig tree (Ficus benjamina), one single individual tree, Banyan garden protected forest, Kenting National Park, Taiwan
Stephen Dalton / Orange tip butterfly (Anthocharis cardamines) male in flight amongst Lady's smock or Cuckooflower (Cardamine pratensis),England.
Mark MacEwen / Angel Falls, the world's highest uninterrupted waterfall with a fall of 807m, flowing from Auyan-tepui, a table-top mountain. Helicopter used in filming at base of waterfall with people beside it....
Erlend Haarberg / White wagtail (Motacilla alba) reflected in water. Pasvik, Norway. May.
Carol Walker / Andalusian horsea, two stallions coming together in arena one dappled grey and one pure white. Spain.
Simon Littlejohn / Bonelli's eagle chicks (Aquila fasciata) in nest pecking each other, aged one week, Andalusia , Spain, May.
Suzi Eszterhas / Northern muriqui monkey (Brachyteles hypoxanthus) juvenile aged one year, in tree playing with its mother's tail, RPPN Feliciano Miguel Abdala, Atlantic Forest, Brazil. June.
Yva Momatiuk & John Eastcott / Adelie Penguin (Pygoscelis adeliae) flapping on iceberg, Antarctic Peninsula, Antarctica
Yva Momatiuk & John Eastcott / Cheetah (Acinonyx jubatus) female yawning, Masai Mara, Kenya
Yva Momatiuk & John Eastcott / Weddell Seal (Leptonychotes weddellii), Antarctic Peninsula, Antarctica
Yva Momatiuk & John Eastcott / Eastern Grey Kangaroo (Macropus giganteus) male joey grazing, Yuraygir National Park, New South Wales, Australia
Yva Momatiuk & John Eastcott / Eastern Grey Kangaroo (Macropus giganteus) yawning, Yuraygir National Park, New South Wales, Australia
Yva Momatiuk & John Eastcott / Gentoo Penguin (Pygoscelis papua) jumping from iceberg, Gerlache Passage, Antarctic Peninsula, Antarctica