Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists, acting as ecosystem engineers by maintaining open meadows and digg...
Andres M. Dominguez / Guillemots (Uria aalge) in flight over snow, Vardo, Norway, March.
Danny Green / Arctic fox (Alopex lagopus) standing in tundra. Svalbard, Norway, July.
Danny Green / Polar bear (Ursus maritimus) male in snow. Svalbard, Norway, April.
Heather Angel / Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China.
Derek Galon / Damaged forest overgrown by various vines, a typical scene in western part of Dominica, West Indies. After hurricane Maria an unprecedented number of vines took advantage of suddenly open spaces in th...
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), four chicks in moult aged 20-24 weeks, parents away fishing in open water. Atka Bay, Antarctica. January.
Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Tui De Roy / Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos.
Guy Edwardes / Hippopotamus (Hippopotamus amphibius) in Chobe River, Botswana.
Guy Edwardes / Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
Andrew Parkinson / Mountain hare (Lepus timidus) three adults on a snow slope high in the Cairngorm Mountains, Cairngorms National Park, Scotland, UK, February. Finalist in the Mammals category of the Wildlife Photogra...
Denis-Huot / Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Alex Hyde / Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio, Cordillera de Talamanca mou...
Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.
Guy Edwardes / View over Pednvounder Beach towards Logan Rock, Porthcurno, Cornwall, England, UK. April 2009.
Floris van Breugel / California poppies (Eschscholzia californica) and California goldfields (Lasthenia californica) extend as far as the eye can see, Antelope Valley Poppy Preserve, California, USA
Bence Mate / Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica.
Wim van den Heever / Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April.
Wim van den Heever / Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015.
Wim van den Heever / Quiver Tree forest, (Aloe dichotoma) at dusk. Keetmanshoop, Namibia.
Eric Baccega / African elephant (Loxodonta africana) and black rhinoceros (Diceros bicornis) bathing at waterhole at night with moon, Okaukuejo, Etosha National Park, Namibia, Africa. Taken with infrared camera.
Ben Hall / Marine Iguana (Amblyrhynchus cristatus) on rock with lava lizard sitting on its head, Galapagos Islands, May.
Danny Green / Red fox (Vulpes vulpes) scratching in the snow, Churchill, Cananda, November
Wim van den Heever / Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June.
Wim van den Heever / Lion (Panthera leo) male in darkness, Okavango Delta, Botswana.
Wim van den Heever / Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.
Jurgen Freund / Split level view of coral reef, Komodo National Park, Flores, Lesser Sunda Islands, Indonesia. Second Place in the Portfolio Award of the Terre Sauvage Nature Images Awards Competition 2015.
Bence Mate / Great egret (Ardea alba) close up of feathers, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.
Bence Mate / Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.
Mark Hamblin / Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015.
Fabrice Cahez / Red fox (Vulpes vulpes) portrait of juvenile, Vosges, France, July.
Nick Garbutt / Darwin's hawkmoth (Xanthopan morgani praedicta) showing extraordinarily long coiled tongue. Andasibe-Mantadia National Park, Madagascar.
Mark Bowler / Black capuchin, (Sapajus nigritus) female standing on hind legs, Rio de Janeiro, Brazil.