Heather Angel / Giant panda (Ailuropoda melanoleuca) portrait, in falling snow. Captive bred, in enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008.
Alex Mustard / Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka.
Tony Wu / Humpback whale (Megaptera novaeangliae australis) male calf playing at ocean surface after coming up to breathe. Vava'u Islands, Tonga.
Staffan Widstrand / African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
John Shaw / James' flamingo / Puna flamingo (Phoenicoparrus jamesi). Lago Colorado, Bolivia. March.
George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.
Ross Hoddinott / Banded demoiselle (Calopteryx splendens) male, resting on grass head, Lower Tamar Lakes, Cornwall, UK. June.
Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.
Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.
Michael Hutchinson / Pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, Devon, England, UK, June.
Michael Hutchinson / Male Marsh fritillary butterfly (Euphydryas aurinia) displaying to a female on a Buttercup (Ranunculus) flower, Devon, England, UK, June.
SCOTLAND: The Big Picture / Puffin (Fratercula artica) with orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June.
Klein & Hubert / Young Hippo (Hippopotamus amphibius) trotting to the Luangwa River at sunrise, Luangwa National Park, Zambia.
John Chan / Male California least tern (Sternula antillarum browni) presenting fish offering to female during mating, Huntington Beach, California, USA, May.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male examining dead chick on ice. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) preparing to jump over crack in sea ice. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), pair mating. Atka Bay, Antarctica. May.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults with chicks aged six to eight weeks. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks near adult's feet. Atka Bay, Antarctica. September.
SCOTLAND: The Big Picture / European badger (Meles meles) foraging in front of house at night. Glenfeshie, Cairngorms National Park, Scotland, UK.
SCOTLAND: The Big Picture / Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK.
Andy Rouse / African Lion (Panthera leo) male at sunrise, Masai Mara, Kenya
Nick Hawkins / Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos.
Nick Hawkins / Galapagos green turtle (Chelonia mydas agassizi) swimming above sea floor with shoal of fish in background. Off San Cristobal Island, Galapagos.
Nick Hawkins / Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica.
Paul Hobson / Red deer (Cervus elaphus) stags in snowy Pine forest. Scotland, UK. March.
Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
Paul Hobson / Rivoli's hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica.
Paul Hobson / Leaf-nosed bat (Phyllostomidae sp), three nectaring on flower. Costa Rica.
Paul Hobson / Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica.
Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Paul Hobson / Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica.
Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
Paul Hobson / Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June.
Paul Hobson / Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October.
Paul Hobson / Badger (Meles meles) crossing road in residential area at night. Sheffield, England, UK. October 2018.
Paul Hobson / Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica.
Markus Varesvuo / Black woodpecker (Dryocopus martius) male perched on tree trunk, trees covered in hoar frost. Kuusamo, Northern Ostrobothnia, Finland. February. Commended in the Birds in the Environment Category of t...
Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
Sven Zacek / Ural owl (Strix uralensis) female and nestling on nest in tree stump. Tartumaa, Southern Estonia. May.
Sven Zacek / Beaver (Castor fiber) felling a Birch (Betula sp) tree for winter supplies. Tartumaa, Southern Estonia. November.
Tui De Roy / Flightless cormorant (Phalacrocorax harrisi), two on rock. Bird drying wings in foreground with other sitting on nest in background. Cape Douglas, Fernandina Island, Galapagos.
Tui De Roy / Blue-footed booby (Sula nebouxii), pair in courtship display. Punta Pitt, San Cristobal Island, Galapagos. April 2017.
Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) walking, Alcedo Volcano, Isabela Island, Galapagos