Lucas Bustamante / Choco Toucan (Ramphastos brevis) on a tree with epiphytes, Mashpi, Pichincha, Ecuador

Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.

Nick Garbutt / Toco toucan (Ramphastos toco) acrobatically perched , feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil.

Hermann Brehm / Toco Toucan (Ramphastos toco) , Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) portrait, Pantanal Brazil.

Angelo Gandolfi / Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil

Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus) calling, captive.

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus), captive.

Nick Garbutt / Toco Toucans (Ramphastos toco) taking flight from the forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Nick Garbutt / Toco Toucan (Ramphastos toco) feeding on fruit in forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Sue Flood / Chestnut mandibled toucan (Ramphastos ambiguus swainsonii) near Boca Tapada, Costa Rica. January 2011

Luiz Claudio Marigo / Channel-billed / Ariel toucan (Ramphastos vittelinus ariel) perched in flowering tree in city street, Rio de Janeiro city, Southeastern Brazil.

Nick Garbutt / A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) attacking a Brown-hooded Parrot (Pionopsitta haematotis) Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) in aggressive encounter with a Montezuma Oropendola (Psarocolius montezuma), Laguna del Lagarto, Santa Rita, Costa Rica

Mary McDonald / Chestnut-mandibled Toucan (Ramphastos ambiguus swainsonii) adult bird perched on the edge of the forest. Laguana del Lagarto, Costa Rica, Central America.

Piper Mackay / Toco toucan (Ramphastos toco) feeding, throwing nut into air to catch it in beak, Pantanal, Brazil.

Edwin Giesbers / Toco toucan (Ramphastos toco), close-up of face, Costa rica

Edwin Giesbers / Keel billed Toucan (Ramphastos sulfuratus), Costa Rica

Luiz Claudio Marigo / Toco toucan {Ramphastos toco} Brazil

Luiz Claudio Marigo / Channel billed toucan {Ramphastos vitellinus} South America

Staffan Widstrand / Toco toucan in tree {Ramphastos toco} Pantanal, Brazil

Nick Gordon / Yellow ridged toucan eating acai palm fruit. Amazonia, Brazil South America

Dietmar Nill / Keel Billed Toucan, Costa Rica.

Neil Lucas / Chestnut mandibled toucan, Panama, Central America

Glenn Bartley/ BIA / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Glenn Bartley/ BIA / Black-mandibled Toucan (Ramphastos ambiguus) during rainfall, Costa Rica

Thomas Marent / Keel-billed Toucan (Ramphastos sulfuratus), native to Central and South America

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Glenn Bartley / Chestnut-mandibled Toucan (Ramphastos swainsonii), Costa Rica

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) portrait, rainforest in Costa Rica

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) perched in tree, Costa Rican rainforest

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) perched in tree, Costa Rican rainforest

Tim Fitzharris / Keel-billed Toucan (Ramphastos sulfuratus) pair, Costa Rica

Kevin Schafer / Red-breasted Toucan (Ramphastos dicolorus) calling, Itatiaia National Park, Brazil

Claus Meyer / Red-breasted Toucan (Ramphastos dicolorus) close-up through vegetation, south Brazil

Claus Meyer / Toco Toucan (Ramphastos toco) calling while perching on branch, Cerrado, Brazil

Pete Oxford / Chestnut-mandibled Toucan (Ramphastos swainsonii), Choco Forest, Ecuador

Gerry Ellis / Keel-billed Toucan (Ramphastos sulfuratus) portrait showing barbed tongue, native to Mexico and Central America

Norbert Wu / Toco Toucan (Ramphastos toco) bill is actually light and spongy, Brazil

Norbert Wu / Toco Toucan (Ramphastos toco) bill is actually light and spongy, Brazil

Nick Garbutt / RF - Black-mandibled toucan (Ramphastos ambiguus) perched on branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)

Nick Garbutt / RF - Keel-billed toucan (Ramphastos sulfuratus) perched on branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)

Nick Garbutt / RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)

Nick Garbutt / Keel-billed toucan (Ramphastos sulfuratus) looking downwards, perched on branch. Boca Tapada, Costa Rica.