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    • Ruff
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Ruffed grouse (Bonasa umbellus) displaying neck "ruff". The ruff is most often displayed once a female is present. Okanogan County, Washington. April.  -  Gerrit  Vyn
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Gerrit Vyn / Ruffed grouse (Bonasa umbellus) displaying neck "ruff". The ruff is most often displayed once a female is present. Okanogan County, Washington. April.
Rear view of Ruff (Philomachus pugnax) in full display at the lek.  Varanger, Finmark, Norway, May.  -  Roger Powell
01450025
Roger Powell / Rear view of Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May.
Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May.  -  Roger Powell
01450013
Roger Powell / Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May.
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June  -  Markus Varesvuo
01412546
Markus Varesvuo / Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Ruffed grouse (Bonasa umbellus) male displaying neck "ruff". The ruff is most often displayed once a female has been attracted by the males drumming and is present. Okanogan County, Washington, USA Ap...  -  Gerrit Vyn
01377469
Gerrit Vyn / Ruffed grouse (Bonasa umbellus) male displaying neck "ruff". The ruff is most often displayed once a female has been attracted by the males drumming and is present. Okanogan County, Washington, USA Ap...
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia  -  Sergey Gorshkov
01375106
Sergey Gorshkov / Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia  -  Sergey Gorshkov
01375096
Sergey Gorshkov / Ruff displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June  -  Loic Poidevin
01361105
Loic Poidevin / Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June
Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June  -  Loic Poidevin
01361104
Loic Poidevin / Ruff (Philomachus pugnax) two rival males displaying, fighting, Varanger, Norway, June
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May  -  Loic Poidevin
01361094
Loic Poidevin / Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
Ruff (Philomachus pugnax) male showing off his ruff in a courtship display behaviour. Varanger, Norway, June.  -  Roger Powell
01338976
Roger Powell / Ruff (Philomachus pugnax) male showing off his ruff in a courtship display behaviour. Varanger, Norway, June.
Ruff (Philomachus pugnax) male in full display. This was the dominant male at the lek. Varanger, Norway, June.  -  Roger Powell
01338968
Roger Powell / Ruff (Philomachus pugnax) male in full display. This was the dominant male at the lek. Varanger, Norway, June.
Two male Ruff (Philomachus pugnax) of different colour types competitively displaying at a lek in the background. Varanger, Norway, June.  -  Roger Powell
01338966
Roger Powell / Two male Ruff (Philomachus pugnax) of different colour types competitively displaying at a lek in the background. Varanger, Norway, June.
Ruff (Philomachus pugnax) male preening, UK, May  -  Ernie Janes
01336706
Ernie Janes / Ruff (Philomachus pugnax) male preening, UK, May
Ruff (Philomachus pugnax) male displaying to attract the attention of a Reeve (female) nearby, Varanger plateau, Arctic Norway. June  -  Roger Powell
01312515
Roger Powell / Ruff (Philomachus pugnax) male displaying to attract the attention of a Reeve (female) nearby, Varanger plateau, Arctic Norway. June
Three male Ruffs (Philomachus pugnax) preparing to display on the communal lekking ground, Varanger plateau, arctic Norway. June  -  Roger Powell
01312508
Roger Powell / Three male Ruffs (Philomachus pugnax) preparing to display on the communal lekking ground, Varanger plateau, arctic Norway. June
Ruff (Philomachus pugnax) male foraging along shore in winter plumage, Lancashire, UK, February  -  Alan Williams
01251800
Alan Williams / Ruff (Philomachus pugnax) male foraging along shore in winter plumage, Lancashire, UK, February
Ruff (Philomachus pugnax) males displaying in rain at lek, Norway  -  Winfried Wisniewski
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Winfried Wisniewski / Ruff (Philomachus pugnax) males displaying in rain at lek, Norway
Ruff (Philomachus pugnax) male displaying at lek, Varanger Peninsula, Norway  -  Winfried Wisniewski
00557169
Winfried Wisniewski / Ruff (Philomachus pugnax) male displaying at lek, Varanger Peninsula, Norway
Andean Condor (Vultur gryphus) adult male raising ruff to keep head warm, Condor Huasi Project, Hacienda Zuleta, Cayambe, Ecuador  -  Tui De Roy
00141138
Tui De Roy / Andean Condor (Vultur gryphus) adult male raising ruff to keep head warm, Condor Huasi Project, Hacienda Zuleta, Cayambe, Ecuador
Ruff (Philomachus pugnax) juvenile feeding whilst on autumn migration. Norfolk, England, UK, August.  -  Oscar Dewhurst
01653024
Oscar Dewhurst / Ruff (Philomachus pugnax) juvenile feeding whilst on autumn migration. Norfolk, England, UK, August.
Common redshank (Tringa totanus) bathing in a shallow freshwater pool, with three courting Ruff (Philomachus pugnax) nearby, Gloucestershire, England, UK, February.  -  Nick Upton
01650724
Nick Upton / Common redshank (Tringa totanus) bathing in a shallow freshwater pool, with three courting Ruff (Philomachus pugnax) nearby, Gloucestershire, England, UK, February.
Mixed flock of Common redshank (Tringa totanus) and Ruff (Philomachus pugnax) resting and preening on a shingle spit, Gloucestershire, England, UK, February.  -  Nick Upton
01650723
Nick Upton / Mixed flock of Common redshank (Tringa totanus) and Ruff (Philomachus pugnax) resting and preening on a shingle spit, Gloucestershire, England, UK, February.
Mixed flock of Common redshank (Tringa totanus), Ruff (Philomachus pugnax) and a Black-headed gull (Chroicocephalus ridibundus) resting and preening on a shingle spit, Gloucestershire, England, UK, Fe...  -  Nick Upton
01650722
Nick Upton / Mixed flock of Common redshank (Tringa totanus), Ruff (Philomachus pugnax) and a Black-headed gull (Chroicocephalus ridibundus) resting and preening on a shingle spit, Gloucestershire, England, UK, Fe...
Ruff (Calidris pugnax) foraging in a lake amongst roosting Northern lapwings (Vanellus vanellus), showing winter plumage, Gloucestershire, England, UK, November.  -  Nick Upton
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Nick Upton / Ruff (Calidris pugnax) foraging in a lake amongst roosting Northern lapwings (Vanellus vanellus), showing winter plumage, Gloucestershire, England, UK, November.
Small flock of Ruff (Philomachus / Calidris pugnax) in flight overhead, Gloucestershire, UK, November.  -  Nick Upton
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Nick Upton / Small flock of Ruff (Philomachus / Calidris pugnax) in flight overhead, Gloucestershire, UK, November.
Ruff (Calidris pugnax) male, Denmark, May  -  Hanne & Jens Eriksen
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Hanne & Jens Eriksen / Ruff (Calidris pugnax) male, Denmark, May
Ruff (Philomachus pugnax) territorial male fighting with satellite males during lek, Varanger, Norway, May  -  Loic Poidevin
01638896
Loic Poidevin / Ruff (Philomachus pugnax) territorial male fighting with satellite males during lek, Varanger, Norway, May
Ruff (Philomachus pugnax) displaying territorial male, Varanger, Norway, May  -  Loic Poidevin
01638895
Loic Poidevin / Ruff (Philomachus pugnax) displaying territorial male, Varanger, Norway, May
Ruff (Philomachus pugnax) male, Varanger, Norway, May  -  Loic Poidevin
01638894
Loic Poidevin / Ruff (Philomachus pugnax) male, Varanger, Norway, May
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May.  -  Loic Poidevin
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Loic Poidevin / Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May.
Ruff (Philomachus pugnax), male displaying at lek. Varanger, Norway. May.  -  Loic Poidevin
01630484
Loic Poidevin / Ruff (Philomachus pugnax), male displaying at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only  -  Loic Poidevin
01630483
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.  -  Loic Poidevin
01630480
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.
Ruff (Philomachus pugnax) three males in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630479
Loic Poidevin / Ruff (Philomachus pugnax) three males in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.  -  Loic Poidevin
01630477
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630475
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display. Varanger, Norway. May.  -  Loic Poidevin
01630467
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630465
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax), two males in competition during courtship display, at lek. Varanger, Norway. May.  -  Loic Poidevin
01630461
Loic Poidevin / Ruff (Philomachus pugnax), two males in competition during courtship display, at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male at lek. Varanger, Norway. May.  -  Loic Poidevin
01630456
Loic Poidevin / Ruff (Philomachus pugnax) male at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male displaying, further male in background. At lek, Varanger, Norway. May.  -  Loic Poidevin
01630454
Loic Poidevin / Ruff (Philomachus pugnax) male displaying, further male in background. At lek, Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630452
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630448
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630447
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630444
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630443
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.  -  Loic Poidevin
01630439
Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.
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