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Ross Hoddinott / Banded demoiselle (Calopteryx splendens) male, resting on grass head, Lower Tamar Lakes, Cornwall, UK. June.
Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.
Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.
Michael Hutchinson / Pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, Devon, England, UK, June.
Michael Hutchinson / Male Marsh fritillary butterfly (Euphydryas aurinia) displaying to a female on a Buttercup (Ranunculus) flower, Devon, England, UK, June.
SCOTLAND: The Big Picture / Puffin (Fratercula artica) with orange sea lichen (Caloplaca marina) covered roch behind, Shiant Isles, Outer Hebrides, Scotland, UK. June.
Klein & Hubert / Young Hippo (Hippopotamus amphibius) trotting to the Luangwa River at sunrise, Luangwa National Park, Zambia.
John Chan / Male California least tern (Sternula antillarum browni) presenting fish offering to female during mating, Huntington Beach, California, USA, May.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male examining dead chick on ice. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) preparing to jump over crack in sea ice. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), pair mating. Atka Bay, Antarctica. May.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults brooding chicks. One chick exhibiting aggression towards other. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two adults with chicks aged six to eight weeks. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks near adult's feet. Atka Bay, Antarctica. September.
SCOTLAND: The Big Picture / European badger (Meles meles) foraging in front of house at night. Glenfeshie, Cairngorms National Park, Scotland, UK.
SCOTLAND: The Big Picture / Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK.
Andy Rouse / African Lion (Panthera leo) male at sunrise, Masai Mara, Kenya
Nick Hawkins / Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos.
Nick Hawkins / Galapagos green turtle (Chelonia mydas agassizi) swimming above sea floor with shoal of fish in background. Off San Cristobal Island, Galapagos.
Nick Hawkins / Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica.
Paul Hobson / Red deer (Cervus elaphus) stags in snowy Pine forest. Scotland, UK. March.
Paul Hobson / Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica.
Paul Hobson / Rivoli's hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica.
Paul Hobson / Leaf-nosed bat (Phyllostomidae sp), three nectaring on flower. Costa Rica.
Paul Hobson / Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica.
Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Paul Hobson / Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica.
Paul Hobson / Red fox (Vulpes vulpes), two cubs play fighting. Sheffield, England, UK. May.
Paul Hobson / Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June.
Paul Hobson / Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October.
Paul Hobson / Badger (Meles meles) crossing road in residential area at night. Sheffield, England, UK. October 2018.
Paul Hobson / Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica.
Pal Hermansen / Brown bear (Ursus arctos), two showing aggression at night. Finland. August.
Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
Sven Zacek / Ural owl (Strix uralensis) female and nestling on nest in tree stump. Tartumaa, Southern Estonia. May.
Sven Zacek / Beaver (Castor fiber) felling a Birch (Betula sp) tree for winter supplies. Tartumaa, Southern Estonia. November.
Tui De Roy / Alcedo giant tortoise (Chelonoidis vandenburghi) walking, Alcedo Volcano, Isabela Island, Galapagos
Pedro Narra / Hedgehog (Erinaceus europaeus) walking through puddle. Sado Estuary, Portugal . March
Steven David Miller / Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia.
Christophe Courteau / Mountain gorilla (Gorilla beringei beringei), dominant silverback in forest clearing. Volcanoes National Park, Rwanda.
Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) blackback, juvenile male demonstrating power. Volcanoes National Park, Rwanda.
Jack Dykinga / Burrowing owl (Athene cunicularia), two huddled together. Marana, Pima County, Arizona, USA.
David Fleetham / Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii.
David Fleetham / Whale shark (Rhincodon typus), a commercial encounter for tourists with feeder above on canoe. Oslob, Cebu, Philippines. April 2017.
David Fleetham / Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines.
David Fleetham / Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii.
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