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    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic
    01397249
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic
    Dave Watts / Wedge-tailed eagle {Aquila audax fleayi} tasmanian sub species, adult portrait, Australia
    01104403
    Dave Watts / Wedge-tailed eagle {Aquila audax fleayi} tasmanian sub species, adult portrait, Australia
    Ben Cranke / Southern elephant seals (Mirounga leonina) subadult males practice sparring. St Andrew's Bay, South Georgia Island
    01681231
    Ben Cranke / Southern elephant seals (Mirounga leonina) subadult males practice sparring. St Andrew's Bay, South Georgia Island
    Jeff Foott / Gorilla (Gorilla beringei beringei) sub adult male feeding on vegetation, close up. Volcanoes National Park, Rwanda, Africa.
    01674979
    Jeff Foott / Gorilla (Gorilla beringei beringei) sub adult male feeding on vegetation, close up. Volcanoes National Park, Rwanda, Africa.
    Tui De Roy / Galapagos sea lions (Zalophus wollebaeki) sub adult males playing with starfish, Mosquera Islet, Galapagos.
    01650988
    Tui De Roy / Galapagos sea lions (Zalophus wollebaeki) sub adult males playing with starfish, Mosquera Islet, Galapagos.
    Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.
    01626301
    Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.
    Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.
    01624122
    Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.
    Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.
    01624121
    Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.
    SCOTLAND: The Big Picture / Mountain hare (Lepus timidus), sub-adult in summer coat on moorland, Scotland, UK, August.
    01623831
    SCOTLAND: The Big Picture / Mountain hare (Lepus timidus), sub-adult in summer coat on moorland, Scotland, UK, August.
    Anup Shah / Lion-tailed macaque (Macaca silenus) sub-adult male sitting in a tree. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.
    01446928
    Anup Shah / Lion-tailed macaque (Macaca silenus) sub-adult male sitting in a tree. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.
    01423099
    Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.
    01423092
    Ammonite / Group of sub-adult African lions (Panthera leo) chasing and jumping on a mother Hippopotamus (Hippopotamus amphibius) and her calf, footage taken at night using infrared camera technology, Masai Mara,...
    01423091
    Ammonite / Sub-adult African lioness (Panthera leo) chasing and jumping onto the back of a Hippopotamus (Hippopotamus amphibius), footage taken at night using infrared camera technology, Masai Mara, Kenya.
    01423090
    Ammonite / Sub-adult African lioness (Panthera leo) watching, chasing and missing a small group of Warthogs (Phacochoerus aethiopicus), Masai Mara, Kenya.
    01421514
    Mark Hamblin / 2020VISION / Two sub-adult Wild Domestic goats (Capra hircus) play fighting on hillside, Cairngorms National Park, Scotland, UK, February.
    Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.
    01417042
    Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.
    Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.
    01417012
    Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male sitting on a beach, Tangkoko National Park, Sulawesi, Indonesia.
    01416902
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male sitting on a beach, Tangkoko National Park, Sulawesi, Indonesia.
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male resting on a fallen tree, Tangkoko National Park, Sulawesi, Indonesia.
    01416835
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male resting on a fallen tree, Tangkoko National Park, Sulawesi, Indonesia.
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male trying to break into bamboo to feed on ant eggs from a nest, Tangkoko National Park, Sulawesi, Indonesia.
    01416821
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male trying to break into bamboo to feed on ant eggs from a nest, Tangkoko National Park, Sulawesi, Indonesia.
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male yawning, Tangkoko National Park, Sulawesi, Indonesia.
    01416817
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male yawning, Tangkoko National Park, Sulawesi, Indonesia.
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male head and shoulders portrait, Tangkoko National Park, Sulawesi, Indonesia.
    01416796
    Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male head and shoulders portrait, Tangkoko National Park, Sulawesi, Indonesia.
    Fiona Rogers / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years standing with sub-adult male 'Kunga' aged 13 years in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Cent...
    01406218
    Fiona Rogers / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years standing with sub-adult male 'Kunga' aged 13 years in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Cent...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...
    01406148
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.
    01406146
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.
    01406145
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. No...
    01406142
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. No...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    01406129
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    01406128
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years looking at his hand, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 20...
    01406127
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years looking at his hand, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 20...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    01406126
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses while juvenile male 'Mobangi' aged 5 years feeds on rotten wood in the background, Bai...
    01406119
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses while juvenile male 'Mobangi' aged 5 years feeds on rotten wood in the background, Bai...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on rotting wood, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...
    01406086
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on rotting wood, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river while sub-adult female 'Mosoko' aged 8 years crosses bi-pedally, Bai Hokou, Dza...
    01406083
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river while sub-adult female 'Mosoko' aged 8 years crosses bi-pedally, Bai Hokou, Dza...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river watched by sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' a...
    01406082
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river watched by sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' a...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years approaching a river while sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 y...
    01406081
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years approaching a river while sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 y...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) rear view of sub-adult male 'Kunga' aged 13 years walking along a fallen tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...
    01406080
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) rear view of sub-adult male 'Kunga' aged 13 years walking along a fallen tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years using her hand to scoop water from a river to drink, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Cen...
    01406067
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years using her hand to scoop water from a river to drink, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Cen...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years peering through branches, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...
    01406060
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years peering through branches, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...
    01406058
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years climbing down a tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2...
    01406057
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years climbing down a tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years playing with sub-adult female 'Mosoko' aged 8 years with juvenile male 'Tembo' aged 4 years watching in the backg...
    01406044
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years playing with sub-adult female 'Mosoko' aged 8 years with juvenile male 'Tembo' aged 4 years watching in the backg...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting on a fallen tree feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afric...
    01406033
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting on a fallen tree feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afric...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...
    01406029
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years drinking from a dirty pool of water in Bai Hokou, Dzanga Sangha Specia...
    01406028
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 years drinking from a dirty pool of water in Bai Hokou, Dzanga Sangha Specia...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...
    01406025
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years pirouetting with excitement, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    01406021
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years pirouetting with excitement, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...
    01406020
    Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years playing with juvenile male 'Tembo' aged 4 years, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central A...
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