Anup Shah / Lion (Panthera leo) male approaching, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.
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Nick Garbutt / Leopard (Panthera pardus) female drinking at night. South Luangwa National Park, Zambia.
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Fabrice Cahez / Wild cat (Felis sylvestris) in snow, Vosges, France, February.
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Mark Bowler / White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru.
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Richard Kirby / Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February.
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Uri Golman / Greenland reindeer (Rangifer tarandus groenlandicus) in velvet, Kangerlussuaq, Greenland, June.
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Sharon Heald / African buffalo (Syncerus caffer) low angle portrait, Chobe National Park, Botswana.
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Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) in urban environment Cardiff, UK
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Gabriel Rojo / Wild puma (Puma concolor) walking across rocks, Torres del Paine National Park, Chile.
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Danny Green / Steller's sea eagle (Haliaeetus pelagicus) close up portrait, Japan, February
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Danny Green / Mountain Hare (Lepus timidus) resting with ears back, Scotland, March. Highly Commended in the Animal Portrait Category of the BWPA Competition 2015.
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Lynn M Stone / Free-range New Hampshire rooster, by corn pile in snow-covered field. Higganum, Connecticut, USA.
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Cyril Ruoso / Komodo dragon (Varanus komodoensis) in habitat with boat, Rinca Island, Komodo National Park, Indonesia. Non-ex
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David Pattyn / Alpine ibex (Capra ibex), portrait of young male. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September.
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Theo Webb / Female Bonobo (Pan paniscus) resting, Max Planck research site LuiKotale in Salonga National Park, Democratic Republic of Congo.
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Alex Hyde / Mountain hare (Lepus timidus) on snow, Cairngorms National Park, Scotland. January.
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Paul Williams / Red leaf monkey (Presbytis rubicunda) mother holding baby, Danum Valley, Sabah, Borneo, Malaysia August.
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Andy Rouse / Little owl (Athene noctua) on old wheel, UK, April.
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Marie Read / American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May.
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Terry Whittaker / Family of Red foxes (Vulpes vulpes) on railway embankment, Kent, UK, July.
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Ben Cranke / Coyote (Canis latrans) in the snow, Yellowstone National Park, Wyoming, USA. January.
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David Pattyn / Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013.
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David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Nature's Best Photography Windland Smith Rice International Awards Competition...
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Igor Shpilenok / Red Fox (Vulpes vulpes) portrait, resting on snow. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March.
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Gerrit Vyn / Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA
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Andy Rouse / African Lion (Panthera leo) adult looking from behind tree, half hidden. South Luangwa, Zambia, Africa.
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Brent Stephenson / Domestic Sheep (Ovis aries) ewe and two lambs, probably Romney x Perendale. Unshorn. Cape Kidnappers, Hawkes Bay, New Zealand, September.
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Andrew Parkinson / Mountain hare (Lepus timidus) alert portrait in white winter coat, Monadhliath Mountains, Scotland, UK, February.
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Matthew Maran / Black Bear (Ursus americanus) with a Chinook Salmon (Oncorhynchus tshawytscha) in its mouth. Ucluth Inlet, Barkley Sound, Vancouver Island, Canada, September.
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Andy Rouse / Long Eared Owl (Asio otus) UK, controlled conditions, April, Captive
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Mark Taylor / Portrait of a Beagle puppy.
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Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...
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Edwin Giesbers / Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June.
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Ernie Janes / Domestic beef cattle in winter rearing shed, Scotland, UK, November 2007
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Edwin Giesbers / Buffy fish owl (Ketupa ketupa) head portrait, captive
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Edwin Giesbers / Orang utan (Pongo pygmaeus) head portrait of dominant male called Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
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Andrew Parkinson / Coot (Fulica atra) portrait, revealing the size of its enormous feet as it crosses a frozen lake. Derbyshire, UK, January (non-ex) Highly commended in the Portfolio category of the BWPA Competition 20...
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Stephen Dalton / Greater kestrel (Falco rupicoloides) perched on telegraph pole, South Africa
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Laurent Geslin / European beaver (Castor fiber) at night, France
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Mark Taylor / Basset Hound puppy, Betty, 9 weeks, walking across.
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Cyril Ruoso / European hare (Lepus europaeus) portrait. Yonne, France. May.
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Jack Dykinga / Burrowing owl (Athene cunicularia) portrait in evening light. Marana, Arizona, USA.
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Terry Whittaker / Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February.
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Konrad Wothe / Papuan boys with King of Saxony bird of paradise (Pteridophora alberti), killed with slingshot. Eastern Highlands, Papua New Guinea. 2019.
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Etienne Littlefair / Northern long-neck turtle (Chelodina rugosa) slowly emerging from hiding in rocks and debris in a small Adelaide River tributary, showing typical daytime wariness of the species, Northern Territory, A...
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David Hall / A young Giant Pacific Octopus (Enteroctopus dofleini) attempting to cross over and avoid touching a row of huge Sunflower sea stars (Pycnopodia helianthoides), Hanson's Island, British Colulmbia,...
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Danny Green / Wolf (Canis lupus) in distance and Grizzly Bear (Ursus arctos) in foreground, Lake Clark National Park, Alaska, September.
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Morley Read / Slow motion clip of an Andean lapwing (Vanellus resplendens) walking in high paramo landscape, looking for predators, Cotopaxi Volcano, Ecuadorian Andes.