Anup Shah / Lion (Panthera leo) male approaching, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Nick Garbutt / Leopard (Panthera pardus) female drinking at night. South Luangwa National Park, Zambia.

Fabrice Cahez / Wild cat (Felis sylvestris) in snow, Vosges, France, February.

Mark Bowler / White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru.

Richard Kirby / Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February.

Uri Golman / Greenland reindeer (Rangifer tarandus groenlandicus) in velvet, Kangerlussuaq, Greenland, June.

Sharon Heald / African buffalo (Syncerus caffer) low angle portrait, Chobe National Park, Botswana.

Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) in urban environment Cardiff, UK

Gabriel Rojo / Wild puma (Puma concolor) walking across rocks, Torres del Paine National Park, Chile.

Danny Green / Steller's sea eagle (Haliaeetus pelagicus) close up portrait, Japan, February

Danny Green / Mountain Hare (Lepus timidus) resting with ears back, Scotland, March. Highly Commended in the Animal Portrait Category of the BWPA Competition 2015.

Lynn M Stone / Free-range New Hampshire rooster, by corn pile in snow-covered field. Higganum, Connecticut, USA.

Cyril Ruoso / Komodo dragon (Varanus komodoensis) in habitat with boat, Rinca Island, Komodo National Park, Indonesia. Non-ex

David Pattyn / Alpine ibex (Capra ibex), portrait of young male. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September.

Theo Webb / Female Bonobo (Pan paniscus) resting, Max Planck research site LuiKotale in Salonga National Park, Democratic Republic of Congo.

Alex Hyde / Mountain hare (Lepus timidus) on snow, Cairngorms National Park, Scotland. January.

Paul Williams / Red leaf monkey (Presbytis rubicunda) mother holding baby, Danum Valley, Sabah, Borneo, Malaysia August.

Andy Rouse / Little owl (Athene noctua) on old wheel, UK, April.

Marie Read / American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May.

Terry Whittaker / Family of Red foxes (Vulpes vulpes) on railway embankment, Kent, UK, July.

Ben Cranke / Coyote (Canis latrans) in the snow, Yellowstone National Park, Wyoming, USA. January.

David Pattyn / Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013.

David Pattyn / Dalmatian Pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece. February. Highly honoured in the Birds Category of Nature's Best Photography Windland Smith Rice International Awards Competition...

Igor Shpilenok / Red Fox (Vulpes vulpes) portrait, resting on snow. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March.

Gerrit Vyn / Spotted owl (Strix nebulosa) Gifford-Pinchot National Forest, Washington USA

Andy Rouse / African Lion (Panthera leo) adult looking from behind tree, half hidden. South Luangwa, Zambia, Africa.

Brent Stephenson / Domestic Sheep (Ovis aries) ewe and two lambs, probably Romney x Perendale. Unshorn. Cape Kidnappers, Hawkes Bay, New Zealand, September.

Andrew Parkinson / Mountain hare (Lepus timidus) alert portrait in white winter coat, Monadhliath Mountains, Scotland, UK, February.

Matthew Maran / Black Bear (Ursus americanus) with a Chinook Salmon (Oncorhynchus tshawytscha) in its mouth. Ucluth Inlet, Barkley Sound, Vancouver Island, Canada, September.

Andy Rouse / Long Eared Owl (Asio otus) UK, controlled conditions, April, Captive

Mark Taylor / Portrait of a Beagle puppy.

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...

Edwin Giesbers / Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June.

Ernie Janes / Domestic beef cattle in winter rearing shed, Scotland, UK, November 2007

Edwin Giesbers / Buffy fish owl (Ketupa ketupa) head portrait, captive

Edwin Giesbers / Orang utan (Pongo pygmaeus) head portrait of dominant male called Richie, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered

Andrew Parkinson / Coot (Fulica atra) portrait, revealing the size of its enormous feet as it crosses a frozen lake. Derbyshire, UK, January (non-ex) Highly commended in the Portfolio category of the BWPA Competition 20...

Stephen Dalton / Greater kestrel (Falco rupicoloides) perched on telegraph pole, South Africa

Laurent Geslin / European beaver (Castor fiber) at night, France

Mark Taylor / Basset Hound puppy, Betty, 9 weeks, walking across.

Cyril Ruoso / European hare (Lepus europaeus) portrait. Yonne, France. May.

Jack Dykinga / Burrowing owl (Athene cunicularia) portrait in evening light. Marana, Arizona, USA.

Terry Whittaker / Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February.

Konrad Wothe / Papuan boys with King of Saxony bird of paradise (Pteridophora alberti), killed with slingshot. Eastern Highlands, Papua New Guinea. 2019.

Etienne Littlefair / Northern long-neck turtle (Chelodina rugosa) slowly emerging from hiding in rocks and debris in a small Adelaide River tributary, showing typical daytime wariness of the species, Northern Territory, A...

David Hall / A young Giant Pacific Octopus (Enteroctopus dofleini) attempting to cross over and avoid touching a row of huge Sunflower sea stars (Pycnopodia helianthoides), Hanson's Island, British Colulmbia,...

Danny Green / Wolf (Canis lupus) in distance and Grizzly Bear (Ursus arctos) in foreground, Lake Clark National Park, Alaska, September.

Morley Read / Slow motion clip of an Andean lapwing (Vanellus resplendens) walking in high paramo landscape, looking for predators, Cotopaxi Volcano, Ecuadorian Andes.