Tony Heald / Hippopotamus {Hippopotamus amphibius} territorial defense at water edge, Chobe national park, Botswana.
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Laurie Hedges / Bison (Bison bison) bull kicking up dust, urinating and then rolling on the ground for scent, Montana, USA, August.
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Marie Read / American bittern (Botaurus lentiginosus) calling / booming in a cattail marsh to mark territory, Salt Alkaline Lake, Kidder County, North Dakota, USA, June.
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Vishal Jadhav / Indian jackal (Canis aureus indicus) pair howling, Maharashtra, India, September.
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Simon Littlejohn / Common kingfisher (Alcedo atthis) perched outside its nest, opening its wings and beak in an aggressive display to a Common blackbird (Turdus merula) which eventually flies away, River Stour, Dorset,...
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Sergey Gorshkov / Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November. Taken with remo...
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David Pattyn / Mute swan (Cygnus olor) in a threathening territorial pose and attacking a rival, Lake Geneva, near Geneva, Switzerland, March
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Oscar Dewhurst / Eurasian coot (Fulica atra) males fighting over territory. London, England, UK. December.;
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Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
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Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
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Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.
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Matthew Maran / Red Fox (Vulpes Vulpes) patrolling the allotment
North London, England, UK, September.
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Andy Rouse / Bengal tiger (Panthera tigris) tigress Arrowhead sniffing tree for scent, Ranthambhore, India
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David Gallan / Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement. Highly Commended 2018 Wildlife Photographer Of The Yea...
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Klein & Hubert / Hippopotamus (Hippopotamus amphibius) charging in Luangwa river, close up of mouth and teeth, Zambia
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Philippe Clement / Territorial Common raven (Corvus corax) on cliff top, tearing out vegetation, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Dave Bevan / Male Pied flycatcher (Ficedula hypoleuca) calling to establish territory and attract a mate, Carmarthenshire, Wales, UK, May.
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Giri Cavale / Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India.
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Jordi Chias / Two male Dusky groupers (Epinephelus marginatus) fighting for territory during mating season at 'baixa do meros', a small pinnacle at Formigas islets Protected area, near Santa Maria Island, Azores. F...
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Jim Manthorpe / Close-up of a Common otter (Lutra lutra) grooming and rolling in seaweed, Western Scotland. March.
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Fred Olivier / Chinstrap penguin (Pygoscelis antarcticus) adding a stick to its nest and fighting with a neighbouring pair, Aitcho Island, South Shetland Islands.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) with head low to the water in aggressive display, The Netherlands. March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Nick Garbutt / Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America.
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David Pattyn / Great crested grebe (Podiceps cristatus) attacking another grebe in the mating season during a territorial dispute. The Netherlands. April.
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David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. April.
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David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. March 2013
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David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. The Netherlands. April 2014
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David Pattyn / Great crested grebe (Podiceps cristatus) adults fighting in a territorial dispute in the mating season. The Netherlands.april 2014
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David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. The Netherlands. April.
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David Pattyn / Great crested grebe (Podiceps cristatus) in a threathening posture trying to intimidate another grebe in a territorial dispute. Head low over the water and crest erected. The Netherlands. April.
01486391
John & Mary-Lou Aitchison / Female European otter (Lutra lutra) on a rock grooming her cub before rolling over and scent marking, Scotland, UK, November.
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Nick Garbutt / Male Bengal tiger (Panthera tigris tigris) known as 'Sundar' (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species.
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Pål Hermansen,Pal Hermansen / Nuthatch (Sitta europaea) chasing away and Black tit (Periparus ater) in car graveyard, Varmland, Sweden, December
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Tui De Roy / Stewart Island Brown Kiwi or Southern Tokoeka (Apteryx australis lawryi), female probing by scent for sand hoppers in rotting kelp beach wrack at night, Ocean Beach, Stewart Island, New Zealand
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Nick Upton / European robin (Erithacus rubecula) standing upright in a territorial threat display in a meadow among patchy snow, Wiltshire, UK, January.
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Shattil & Rozinski,Shattil & Rozinski / Black bear (Ursus americanus) cub climbing a fence, Minnesota, USA, May.
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Denis-Huot,Denis-Huot / Lion (Panthera leo) males feeding on buffalo (Syncerus caffer) with other buffalos watching. Masai Mara, Kenya, Vulnerable species.
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Denis-Huot,Denis-Huot / Lion (Panthera leo) male chasing White backed vultures (Gyps africanus) off its prey, Masai-Mara Game Reserve, Kenya. Vulnerable species.
01426970
Igor Shpilenok,Igor Shpilenok / Red Foxes (Vulpes vulpes) fighting outside ranger cabin window. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, February.
01425281
Denis-Huot / African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
01424764
Andy Trowbridge / Great Tit (Parus major) perched on frost covered spruce branch with wings open in a show of aggression. Southern Norway, October.
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Paul Hobson / Arctic tern (Sterna paradisaea) two fighting in the air, Scotland, UK July
01412867
Anup Shah / Topi (Damaliscus lunatus jimela) two males sparring, with another approaching, Masai Mara National Reserve, Kenya. March
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Markus Varesvuo / Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
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Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years walking at the edge of the forest, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afr...
