Konrad Wothe / Two male Lawes' parotia birds of paradise (Parotia lawesii) hopping on its display ground, with females watching nearby, Papua New Guinea.

Fred Olivier / Close-up shot tracking down a meltwater waterfall emerging from a glacier, Brasvellbreen Glacier, Magdalena Fjord, Svalbard, Norway, 2016.

John Waters / Large group of Pied wagtails (Motacilla alba) roosting in a Plane tree (Platanus hispanica), Bath, England, UK, January.

Michael Hutchinson / Close-up of a pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, shot tilt downs, Devon, England, UK, June.

Nick Upton / Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.

Steve Downer / Red fox (Vulpes vulpes) jumping over a fence and taking food left out on a tree stump in a garden, Birmingham, England, UK, April.

David Perpinan / Close-up of a Giant anteater (Myrmecophaga tridactyla) eating ants, Pantanal, Mato Grosso do Sul, Brazil.

James Dunbar / Close up of a Cryptic mantis (Sibylla pretiosia) catching and eating a fruit fly, UK. Captive.

Fred Olivier / Close-up of a Black browed albatross (Thalassarche melanophris) preening its chick, chick looks up panting, thermoregulating, West Point Island, Falkland Islands.

Fred Olivier / Tilt shot of a Black browed albatross (Thalassarche melanophris) with its chick on a nest, New Island, Falkland Islands.

Jabruson Motion / Juvenile Eastern lowland gorilla (Gorilla beringei graueri) looking up towards the canopy, Kahuzi-Biega National Park, South Kivu, Democratic Republic of Congo.

Fred Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Two King penguins (Aptenodytes patagonicus) walking along the shore, shot tilts up from feet to head, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.

John & Mary-Lou Aitchison / Flock of Barnacle geese (Branta leucopsis) flying over the sea, Islay, Inner Hebrides, Scotland, UK, November.

Jabruson Motion / Close-up of a young Western gorilla (Gorilla gorilla) 'Tembo', member of the 'Makumba' group, breaking open a termite nest and eating termites, Bai Hokou, Dzanga-Ndoki National Park, Sangha-Mbaere Pre...

Ammonite / Close-up shot panning up a swarm of Honey bees (Apis mellifera) hanging from a fir tree, France, April.

Alex Mustard / Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...

Paul D Stewart / Illustration of 'The Entomologist'. Rare copper print with original hand colouring, drawn by George Spratt, printed by George Madely, and sold individually as part of a series through Charles Tilt's F...

Neil Aldridge / A tilting up shot revealing the Lesser butterfly orchid (Platanthera bifolia) on moorland, Devon, United Kingdom.

Neil Aldridge / A tilt shot revealing a Common spotted orchid (Dactylorhiza fuchsii) in woodland, North Somerset, United Kingdom, June.

Neil Aldridge / A tilt shot revealing a Broad-leaved helleborine (Epipactis helleborine) in ancient woodland, North Somerset, United Kingdom, July.

Neil Aldridge / A tilt shot revealing a Broad-leaved helleborine (Epipactis helleborine) in ancient woodland, North Somerset, United Kingdom, July.

Konrad Wothe / Male Lawes' parotia bird of paradise (Parotia lawesii) displaying in a rainforest, Papua New Guinea.

Konrad Wothe / Male Lawes' parotia bird of paradise (Parotia lawesii) preening in a rainforest, Papua New Guinea.

Konrad Wothe / Three Male Lawes' parotia birds of paradise (Parotia lawesii) hopping on display ground in a rainforest, Papua New Guinea.

Nick Upton / Tilt shot of a female Wasp spider (Argiope bruennichi) on her web, showing zigzag stabilimenta threads, Wiltshire, England, UK, July.

Laurie Hedges / Tilt shot up a large rainforest tree, Corcovado National Park, Costa Rica, 2019.

Laurie Hedges / Tilt shot up to rainforest canopy, Corcovado National Park, Costa Rica, 2019.

Fred Olivier / Wide-angle shot tilting up from brash ice to glacier and mountain, Magdalena Fjord, Svalbard, Norway, 2016.

Fred Olivier / Wide-angle shot tilting up from brash ice to glacier and mountain, Magdalena Fjord, Svalbard, Norway, 2016.

Fred Olivier / Tilt shot up a glacier face, Magdalena Fjord, Svalbard, Norway, 2016.

Fred Olivier / Close-up of a meltwater waterfall emerging from a glacier, Brasvellbreen Glacier, Magdalena Fjord, Svalbard, Norway, 2016.

Fred Olivier / Close-up of a meltwater waterfall emerging from a glacier, Brasvellbreen Glacier, Magdalena Fjord, Svalbard, Norway, 2016.

Fred Olivier / Close-up of Brasvellbreen Glacier, with meltwater waterfall, tilts down, Brasvellbreen, Svalbard, Norway, 2016.

Fred Olivier / Tilt shot down the terminus of Brasvellbreen Glacier, with meltwater waterfall, Svalbard, Norway, 2016.

Fred Olivier / Tilt shot up from reflections to cliffs, water surface strewn with fragments of glacial ice, near Sermeq glacier, Prince Christian Sound, Greenland, 2016.

Fred Olivier / Tracking shot of reflections in calm fjord, near Sermeq Glacier, Prince Christian Sound, Greenland, 2016.

Fred Olivier / Tilt shot from reflections to Sermeq Glacier, Prince Christian Sound, Greenland, 2016.

Fred Olivier / Retreating glacier, Skoldungen Fjord, Greenland, 2016.

Fred Olivier / Retreating glacier, shot tilts down from glacier tongue to waterfall to dry valley, Greenland, 2016.

Fred Olivier / Wide angle shot of a Polar bear (Ursus maritimus) walking on melting pack ice, Svalbard, Norway, 2016.

Fred Olivier / Sunset over pack ice, Antarctica, 2017.

Fred Olivier / Tilt shot up a Giraffe (Giraffa camelopardalis) with a Yellow billed oxpecker (Buphagus africanus) perched on its back, Serengeti National Park, Tanzania.

Fred Olivier / Tilt-shot up a juvenile Giraffe (Giraffa camelopardalis), Serengeti National Park, Tanzania.

Fred Olivier / Tracking tilt shot of an eroded coral platform, forming a rugged coast, Aniwa island, Vanuatu, 2018.

Morley Read / Path leading into tropical rainforest, Amazon rainforest, Pastaza Province, Ecuador.

Morley Read / Aerial tilt shot over rainforest canopy on a tepui, Rio Nangaritza Valley, Cordillera del Condor. Zamora Chinchipe Province, Ecuador, 2018.