John Waters / Large group of Pied wagtails (Motacilla alba) roosting in a Plane tree (Platanus hispanica), Bath, England, UK, January.

Michael Hutchinson / Close-up of a pair of Marsh fritillary butterflies (Euphydryas aurinia) mating, shot tilt downs, Devon, England, UK, June.

Nick Upton / Tilt shot of a dam constructed by Eurasian beavers (Castor fiber) on a section of the River Otter, Devon, England, UK, June 2018.

Steve Downer / Red fox (Vulpes vulpes) jumping over a fence and taking food left out on a tree stump in a garden, Birmingham, England, UK, April.

David Perpinan / Close-up of a Giant anteater (Myrmecophaga tridactyla) eating ants, Pantanal, Mato Grosso do Sul, Brazil.

James Dunbar / Close up of a Cryptic mantis (Sibylla pretiosia) catching and eating a fruit fly, UK. Captive.

Fred Olivier / Close-up of a Black browed albatross (Thalassarche melanophris) preening its chick, chick looks up panting, thermoregulating, West Point Island, Falkland Islands.

Fred Olivier / Tilt shot of a Black browed albatross (Thalassarche melanophris) with its chick on a nest, New Island, Falkland Islands.

Fred Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Two King penguins (Aptenodytes patagonicus) walking along the shore, shot tilts up from feet to head, Macquarie Island, Sub-Antarctic Australia.

Fred Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.

John & Mary-Lou Aitchison / Flock of Barnacle geese (Branta leucopsis) flying over the sea, Islay, Inner Hebrides, Scotland, UK, November.

Jabruson Motion / Close-up of a young Western gorilla (Gorilla gorilla) 'Tembo', member of the 'Makumba' group, breaking open a termite nest and eating termites, Bai Hokou, Dzanga-Ndoki National Park, Sangha-Mbaere Pre...

Ammonite / Close-up shot panning up a swarm of Honey bees (Apis mellifera) hanging from a fir tree, France, April.

Alex Mustard,Alex Mustard / Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...

Paul D Stewart / Illustration of 'The Entomologist'. Rare copper print with original hand colouring, drawn by George Spratt, printed by George Madely, and sold individually as part of a series through Charles Tilt's F...

Laurie Hedges / Tilt shot up a large rainforest tree, Corcovado National Park, Costa Rica, 2019.

Laurie Hedges / Tilt shot up to rainforest canopy, Corcovado National Park, Costa Rica, 2019.

Morley Read / Path leading into tropical rainforest, Amazon rainforest, Pastaza Province, Ecuador.

Morley Read / Aerial tilt shot over rainforest canopy on a tepui, Rio Nangaritza Valley, Cordillera del Condor. Zamora Chinchipe Province, Ecuador, 2018.

Morley Read / Aerial tilt shot of a waterfall surrounded by rainforest, Amazonian slopes of the Andes, Morona Santiago Province, Ecuador, 2018.

Morley Read / Aerial tilt shot over montane rainforest canopy, Los Cedros Biological Reserve, Imbabura Province, Ecuador, 2017.

Morley Read / Slow motion shot of a mountain stream running through pristine montane rainforest, Carchi Province, Ecuador, 2017.

Michael Hutchinson / Tilt shot up to a pair of Marsh fritillary butterflies (Euphydryas aurinia) mating in grass, Devon, England, UK, June.

Jabruson Motion / Infant Sumatran orangutan (Pongo abelii) separated from mother and isolated in tree for capture, holding a branch and waving it at conservation workers below, part of a relocation programme, Sei Serda...

Jabruson Motion / Infant Sumatran orangutan (Pongo abelii) separated from mother and isolated in tree for capture, holding a branch and waving it at conservation workers below, part of a relocation programme, Sei Serda...

Kerstin Hinze / Red deer (Cervus elaphus) stag yawning and resting during rut, September. Captive.

Philippe Clement / Tilt shot of a frozen lake with reeds, Belgium, February.

Sandesh Kadur / Tilt shot of a pair of Blue footed boobies (Sula nebouxii), displaying, pans up from feet, Galapagos Islands, Ecuador.

Five Films / Tilt shot up a bush growing in a reedbed, with sun setting behind and illuminating reeds, Somerset Levels, England, UK, November.

David Perpinan / Great rufous woodcreeper (Xiphocolaptes major) arriving at dead palm tree containing its nest, Pantanal, Mato Grosso do Sul, Brazil.

David Perpinan / Tilt shot up to a Capybara (Hydrochoerus hydrochaeris), Pantanal, Mato Grosso do Sul, Brazil.

Michael Hutchinson / Tilt shot from Common bluebells (Hyacinthoides non-scripta) flowering in woodland up to the canopy, Prior's Wood Avon Wildlife Trust Reseve, Bristol, England, UK, May.

John Waters / Tilt shot of birdwatchers watching a murmuration of Common starlings (Sturnus vulgaris), Ham Wall RSPB Reserve, Somerset Levels, England, UK, December.

John Waters / Glossy ibis (Plegadis falcinellus) feeding at dawn, Ham Wall RSPB Reserve, Somerset Levels, England, UK, December.

John Waters / Birdwatchers waiting to watch a murmuration of Common starlings (Sturnus vulgaris) at dusk, Ham Wall RSPB Reserve, Somerset Levels, England, UK, December.

John Waters / Wide angle shot of a murmuration of Common starlings (Sturnus vulgaris) over reedbeds at dusk, Ham Wall RSPB Reserve, Somerset Levels, UK, December.

Fred Olivier / Tilt shot of an Emperor penguin (Aptenodytes forsteri) walking, Adelie Land, Antarctica, January.

Fred Olivier / Tilt shot up to a group of Emperor penguin (Aptenodytes forsteri) chicks preening, Adelie Land, Antarctica, January.

Fred Olivier / Tilt shot of an Emperor penguin (Aptenodytes forsteri) chick feeding, Adelie Land, Antarctica, January.

Fred Olivier / Tilt shot of a nearly fledged Emperor penguin (Aptenodytes forsteri) chick, with only juvenile head down feathers remaining, Adelie Land, Antarctica, January.

Fred Olivier / Emperor penguin (Aptenodytes forsteri) chick walking to the sea, following some Adelie penguins (Pygoscelis adeliae), shot tilts up to reveal more chicks crossing a tide crack, Adelie Land, Antarctica...

Fred Olivier / Close-up of a male Adelie penguin (Pygoscelis adeliae) displaying, Adelie Land, Antarctica, January.

Fred Olivier / Close-up of an Adelie penguin (Pygoscelis adeliae) feeding chicks, Adelie Land, Antarctica, January.

Fred Olivier / Male Adelie penguin (Pygoscelis adeliae) preening in colony, Adelie Land, Antarctica, January.

Fred Olivier / Adelie penguins (Pygoscelis adeliae) bathing, Adelie Land, Antarctica, January.

Fred Olivier / Close-up of Adelie penguins (Pygoscelis adeliae) on eggs, Adelie Land, Antarctica, January.

Fred Olivier / Adelie penguins (Pygoscelis adeliae) sitting back on its nest after displaying, Adelie Land, Antarctica, January.