Lucas Bustamante / Choco Toucan (Ramphastos brevis) on a tree with epiphytes, Mashpi, Pichincha, Ecuador

John Shaw / Chestnut-eared aracari (Pteroglossus castanotis) on cat-gut vine. Pantanal, Brazil, August.

David Pattyn / Plate-billed Mountain-Toucan (Andigena laminirostris), Bellavista private reserve, Mindo cloud forest, Pichincha, Ecuador, July

Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.

Nick Garbutt / Toco toucan (Ramphastos toco) acrobatically perched , feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil.

Bence Mate / Collared aracari (Pteroglossus torquatus) squabbling with Montezuma oropendola (Psarocolius montezuma) Costa Rica.

Bence Mate / Collared aracari (Pteroglossus torquatus) two perched, possibly courtship feeding, Costa Rica.

Hermann Brehm / Toco Toucan (Ramphastos toco) , Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) portrait, Pantanal Brazil.

Angelo Gandolfi / Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil

Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus) calling, captive.

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus), captive.

Nick Garbutt / Toco Toucans (Ramphastos toco) taking flight from the forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Nick Garbutt / Toco Toucan (Ramphastos toco) feeding on fruit in forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Sue Flood / Chestnut mandibled toucan (Ramphastos ambiguus swainsonii) near Boca Tapada, Costa Rica. January 2011

Luiz Claudio Marigo / Channel-billed / Ariel toucan (Ramphastos vittelinus ariel) perched in flowering tree in city street, Rio de Janeiro city, Southeastern Brazil.

Nick Garbutt / A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) attacking a Brown-hooded Parrot (Pionopsitta haematotis) Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) in aggressive encounter with a Montezuma Oropendola (Psarocolius montezuma), Laguna del Lagarto, Santa Rita, Costa Rica

Mary McDonald / Chestnut-mandibled Toucan (Ramphastos ambiguus swainsonii) adult bird perched on the edge of the forest. Laguana del Lagarto, Costa Rica, Central America.

Piper Mackay / Toco toucan (Ramphastos toco) feeding, throwing nut into air to catch it in beak, Pantanal, Brazil.

Edwin Giesbers / Toco toucan (Ramphastos toco), close-up of face, Costa rica

Edwin Giesbers / Keel billed Toucan (Ramphastos sulfuratus), Costa Rica

Luiz Claudio Marigo / Toco toucan {Ramphastos toco} Brazil

Luiz Claudio Marigo / Channel billed toucan {Ramphastos vitellinus} South America

Staffan Widstrand / Toco toucan in tree {Ramphastos toco} Pantanal, Brazil

Nick Gordon / Yellow ridged toucan eating acai palm fruit. Amazonia, Brazil South America

Dietmar Nill / Keel Billed Toucan, Costa Rica.

Neil Lucas / Chestnut mandibled toucan, Panama, Central America

Daniel Brockner/ BIA / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Mindo, Ecuador

Glenn Bartley/ BIA / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Glenn Bartley/ BIA / Black-mandibled Toucan (Ramphastos ambiguus) during rainfall, Costa Rica

Thomas Marent / Keel-billed Toucan (Ramphastos sulfuratus), native to Central and South America

Thomas Marent / Red-headed Barbet (Eubucco bourcierii) male, Valle del Cauca, Colombia

Chien Lee / Plate-billed Mountain-Toucan (Andigena laminirostris), Mindo, Ecuador

Chien Lee / Plate-billed Mountain-Toucan (Andigena laminirostris), Mindo, Ecuador

Glenn Bartley/ BIA / Versicolored Barbet (Eubucco versicolor) male, Amazon, Peru

Glenn Bartley/ BIA / Collared Aracari (Pteroglossus torquatus) during rainfall, Costa Rica

Glenn Bartley/ BIA / Saffron Toucanet (Baillonius bailloni), Atlantic Rainforest, Brazil

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius), Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius), Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius) trio, Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Tui De Roy / Plate-billed Mountain-Toucan (Andigena laminirostris) at nest cavity, Bellavista Cloud Forest Reserve, Tandayapa Valley, Andes, Ecuador

Tui De Roy / Plate-billed Mountain-Toucan (Andigena laminirostris), Andes, Ecuador