Andy Rouse / Maasai Giraffe (Giraffa camelopardalis tippelskirchi) silhouetted by beams of light from setting sun, Maasai Mara, Kenya, Africa. Winner of the African section of the Nature’s Best Windland Smith Rice...
Vincent Grafhorst / Minden / Crater of active volcano at twilight, Erta Ale, Danakil Depression, Ethiopia
Pal Hermansen / Oystercatcher (Haematopus ostralegus) landing on coast in midnight sun. Lofoten, Norway. July.
Orsolya Haarberg / View over Mt Saksa, Storebjorn and Kniven in the Jotunheimen National Park in evening light, Norway. April 2020.
Erlend Haarberg / Photographers camping in the Jotunheimen mountains. Mt Kyrkja in the background. Norway. April 2020.
David Hall / Split level of Sockeye salmon (Oncorhynchus nerka) during migration; Adams River, British Columbia, Canada. October.
Ross Hoddinott / Snowdrops (Galanthus nivalis) silhouetted at twilight, double exposure, Broxwater, Cornwall, UK. February 2017.
John Chan / Great horned owl (Bubo virginianus) chick watching for parents to return with food, Southern California, USA, June.
John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.
John Chan / Peregrine falcon (Falco peregrinus) plucking Dowitcher (Limnodromus) prey, Southern California, USA.
John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019
Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) spyhopping, with a snowy landscape in the background, Skjervoy, Troms, Norway, December.
Theo Bosboom / Blue hour before sunrise over the dunes of Seilebost beach, Isle of Harris, Hebrides, Scotland. February 2018.
Jack Dykinga / Backlit silver cholla cactus (Cylindropuntia echinocarpa) surrounded by California poppies (Eschscholzia californica), beneath the eroded boulders that define the park. Joshua Tree National Park, Moja...
Sven Zacek / Eurasian pygmy owl (Glaucidium passerinum) perched on snow covered branch in evening light. Vorumaa, Southern Estonia. November.
Sven Zacek / Mist over bog with conifers in pre-dawn light. Endla Nature Reserve, Jogevamaa, Central Estonia. June 2016.
Sven Zacek / Ural owl (Strix uralensis) female in flight having taken off from nest, in evening light. Tartumaa, Southern Estonia. May.
Sven Zacek / Ural owl (Strix uralensis) female flying with prey in beak. Flying back to nest in evening light. Tartumaa, Southern Estonia. May.
Sven Zacek / Dandelion (Taraxacum officinale), a field of many seedheads highlighted at sunset. Karula National Park, Valgamaa, Southern Estonia. June 2017.
Sven Zacek / Ural owl (Strix uralensis) perched on Alder (Alnus sp) tree at dusk. Tartumaa, Southern Estonia. April.
Nayan Khanolkar / Bengal tiger (Panthera tigris tigris) in forest clearing. Camera trap image. Pench National Park, Madhya Pradesh, India. January 2018.
Guy Edwardes / Gaada Stack, Foula, Shetland, Scotland, UK. August.
Guy Edwardes / Butt of Lewis lighthouse, Port Nis / Port of Ness, Isle of Lewis, Outer Hebrides, Scotland, UK. March, 2015.
Guy Edwardes / Coastal cottages at twilight, Portree, Isle of Skye, Inner Hebrides, Scotland, UK. January 2014.
Guy Edwardes / Sea stacks, Mangurstadh, Aird Feinis, Isle of Lewis, Outer Hebrides, Scotland, UK. March 2014.
Guy Edwardes / Eilean Glas Lighthouse, Isle of Scalpay, Outer Hebrides, Scotland, UK. March 2015.
Guy Edwardes / View across Loch Duich to Eilean Donan Castle, Kyle of Lochalsh, Highlands, Scotland, Uk. July 2012.
Guy Edwardes / Rock formations at Kimmeridge Bay, Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
Guy Edwardes / Rock formations at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.
Theo Bosboom / Emerald damselfly (Lestes sponsa) in habitat, at twilight, Hondenven, Tubbergen, the Netherlands, August.
Theo Bosboom / Emerald damselfly (Lestes sponsa) in habitat in early morning twilight, Hondenven, Tubbergen, the Netherlands, August.
Theo Bosboom / Banded demoiselles (Calopteryx splendens) males roosting among common nettles, Roosendaal, the Netherlands. July.
Theo Bosboom / Blue damselflies (Coenagrionidae) resting in heather flowers in dawn twilight, Hatertse vennen near Nijmegen, the Netherlands, August.
Sergey Gorshkov / Moonrise at twilight, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Michael Hutchinson / European hedgehog (Erinaceus europaeus) in garden at dusk, with house in the background, Bristol, England, UK, 2016.
Theo Bosboom / Limpets (Patellidae) exposed at low tide on the coast of Playa de Ursa, Portugal. Finalist in the Animals in their Environment Category of the Wildlife Photographer of the Year Awards (WPOY) Competiti...
Fred Olivier / Adelie penguins (Pygoscelis adeliae) walking in front of a blue iceberg, Adelie Land, Antarctica, January.
Jack Dykinga / Massive wildflower display with Lanceleaf monolopia 1+Monolopia lanceolata+2, and Orange fiddle neck 1+Amsinckia intermedia+2 The Temblor Range, also carpeted in flowers, in evening light in the back...
Philippe Clement / Flock of Jackdaws (Corvus monedula) congregating at dusk at communal roost, Belgium, December.
Danny Green / RF - Red fox (Vulpes vulpes) in dappled light on snow, Yellowstone National Park, USA, February 2016 (This image may be licensed either as rights managed or royalty free.)
Espen Bergersen / Humpback whales (Megaptera novaeangliae) bubble net / lunge feeding on herring (Clupea harengus) at dusk, pod co-operative feeding, Kvaloya, Troms, Norway, November
Espen Bergersen / Killer whale / Orca (Orcinus orca) male surfacing at sunset, Kvaloya, Troms, Norway October
Connor Stefanison / Common loon (Gavia immer) sits on her nest at sunset in the Cariboo region of British Columbia, Canada, taken with remote, May
David Tipling / Wandering albatross (Diomedea exulans) a pair displaying at dusk, Albatross Island, South Georgia January
David Tipling / Wandering albatross (Diomedea exulans) pair displaying at dusk in ritual bonding behaviour, Albatross Island in Bay of Isles, South Georgia January
Neil Aldridge / Two Impalas (Aepyceros melampus) ewes grooming each other in evening light on South Africa's Kruger National Park.
David Pattyn / Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June