Markus Varesvuo / Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March.
Markus Varesvuo / Black Woodpecker male (Dryocopus martius) Finland, February.
Markus Varesvuo / Sparrowhawk (Accipiter nisus) adult female, Norway, October.
Espen Bergersen / Northern fulmar (Fulmarus glacialis) in front of the Seljalandsfoss waterfall, Iceland. May
Espen Bergersen / Common Loon (Gavia immer) flapping wings in water, Iceland, May.
Andres M. Dominguez / Black-tailed godwit (Limosa limosa) Danube Delta, Romania. July
Andres M. Dominguez / Common firecrest (Regulus ignicapilla) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
Andres M. Dominguez / Common redstart (Phoenicurus phoenicurus) taking off, Sierra de Grazalema Natural Park. Southern Spain. September
Hermann Brehm / Cuckoo (Cuculus canorus) in flight, Germany, April. May
Guy Edwardes / Barn owl (Tyto alba) in flight above post and wildflowers, in morning light. The Hawk Conservancy Trust, Hampshire, England, UK. July. Captive.
George Sanker / Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July.
Piotr Naskrecki / Termite (Macrotermes mossambicus) winged individual on pile of wings shed by other termites, with winged male Carpenter ants (Camponotus). Gorongosa National Park, Mozambique. Highly commended in the...
Nick Upton / Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019.
John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.
John Cancalosi / Great horned owl (Bubo virginianus) chick stretching its wings in nest in a Saguaro (Carnegiea gigantea), is groomed by its parent, Sonoran Desert, Arizona, USA, May.
Oscar Dewhurst / Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
Phil Savoie / Blue tit (Cyanistes caeruleus) bringing Crab spider (Misumena vatia) prey to nest, Monmouthshire Wales, UK, May.
Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.
Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Marie Read / Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her and crossing his bill with hers...
Marie Read / Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April.
Marie Read / Roseate spoonbill (Ajaia ajaja) adult in breeding plumage preening tail feathers, Black Point Wildlife Drive, Merritt Island National Wildlife Refuge, Florida, USA, January.
Marie Read / Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August.
Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Bernard Castelein / Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Bernard Castelein / Darter ( Anhinga melanogaster) drying wings, Keoladeo NP, Bharatpur, India
Bernard Castelein / Intermediate egret (Egrettta intermedia), Keoladeo NP, Bharatpur, India
Andy Rouse / Barn owl (Tyto alba) hunting with sunset light on wings, UK. March.
Alex Hyde / Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November.
David Pattyn / Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Russell Cooper / Mandarin duck drake (Aix galericulata) from behind flapping its wings. Southwest London, UK. February.
Staffan Widstrand / Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January.
George Sanker / Arctic tern (Sterna paradisaea), two fighting in mid-air. Machias Seal Island, New Brunswick / Maine, Canada / USA. July.
John Chan / Elegant tern (Thalasseus elegans) with a broken wing, attempting to take off, Southern California, USA, April.
John Chan / Western gulls (Larus occidentalis) attacking an Elegant tern (Thalasseus elegans) with a broken wing, Southern California, USA, April.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chicks aged 20-24 weeks stretching wings, in moult whilst parents fish at sea. Atka Bay, Antarctica. January.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
SCOTLAND: The Big Picture / Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK.
Paul Hobson / Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK.
Paul Hobson / Rivoli's hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica.
Markus Varesvuo / Blue tit (Parus caeruleus) perched on tree trunk, looking at camera with wing stretched. Haukipudas, Finland. January.