Ann & Steve Toon / RF - Lion (Panthera leo) head portrait at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / Cheetah (Acinonyx jubatus) walking, silhouetted at dusk. Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana.

Ann & Steve Toon / RF - African elephants (Loxodonta africana) in water, trunks touching, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / Ostrich (Struthio camelus) silhouetted in dust storm. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa.

Ann & Steve Toon / Meerkats (Suricata suricatta) on the move, Kgalagadi Transfrontier Park, Northern Cape, South Africa.. Second place in the Mammals Category of the GDT European Nature Photographer of the Year Competi...

Ann & Steve Toon / Photographer Ann Toon photographing Cape ground squirrels (Xerus inauris) Kgalagadi Transfrontier Park, South Africa, January 2016. Model released.

Ann & Steve Toon / Aardvark (Orycteropus afer) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. Medium repro only

Ann & Steve Toon / Spotted eagle owl (Bubo africanus) subadult at nigh by water, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Leopard / mountain tortoise (Stigmochelys pardalis) with elephant bull (Loxodonta africana) behind in water, Addo elephant national park, Eastern Cape, South Africa, September

Ann & Steve Toon / Lion (Panthera leo) male snarling at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Giraffe (Giraffa camelopardalis) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa August

Ann & Steve Toon / Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August

Ann & Steve Toon / Malachite sunbird (Nectarinia famosa) feding at flower, Kirstenbosch National Botanical Garden, Cape Town, South Africa, September

Ann & Steve Toon / Orange-breasted sunbird (Anthobaphes violacea), Kirstenbosch National Botanical Garden, Cape Town, South Africa, September

Ann & Steve Toon / Nile crocodile (Crocodylus niloticus) at night, Zimanga private game reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa.

Ann & Steve Toon / RF - Lion (Panthera leo) male running in desert, Kgalagadi Transfrontier Park, South Africa. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / Cape sugarbird (Promerops cafer) among yellow Pincushion (Leucospermum cordifolium) flowers, Kirstenbosch Botanical Gardens, Cape Town, South Africa, September.

Ann & Steve Toon / Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January.

Ann & Steve Toon / Meerkat (Suricata suricatta) running, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January.

Ann & Steve Toon / Meerkats (Suricata suricatta) with young, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January.

Ann & Steve Toon / White rhino (Ceratotherium simum) at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / White rhino (Ceratotherium simum) at waterhole drinking at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Elephants (Loxodonta africana) at waterhole drinking at night. One spraying water from trunk, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Elephants (Loxodonta africana) at waterhole drinking at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Impala (Aepyceros melampus) with redbilled oxpecker (Buphagus erythrorhynchus), Kruger National Park, South Africa.

Ann & Steve Toon / Pied crows (Corvus albus) perched on horns of antelope skull. Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa.

Ann & Steve Toon / African fish eagle (Haliaeetus vocifer) flying, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa.

Ann & Steve Toon / Arctic tern (Sterna paradisaea) with sandeel. Inner Farne, Northumberland, UK. July.

Ann & Steve Toon / Egyptian goose (Alopochen aegyptiaca) with goslings, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Common wildebeest (Connochaetes taurinus) with reflection at waterhole, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Sable (Hippotragus niger), Chobe National Park, Botswana, June

Ann & Steve Toon / Cape buffalo (Syncerus caffer) at waterhole at night with plains zebra (Equus quagga), Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Cape buffalo (Syncerus caffer) drinking at waterhole at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Plains zebra (Equus quagga) drinking at waterhole at night, Zimanga private game reserve, KwaZulu-Natal, South Africa, September

Ann & Steve Toon / Yellow mongoose (Cynictis penicillata), Kgalagadi transfrontier park, South Africa, June

Ann & Steve Toon / RF - Warthog male (Phacochoerus africanus) drinking, Zimanga game reserve, KwaZulu-Natal, South Africa, June. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / RF - Leopard female (Panthera pardus) watched by two Gemsbok (Oryx gazella), Kgalagadi transfrontier park, South Africa, June. (This image may be licensed either as rights managed or royalty free.)

Ann & Steve Toon / Ostrich (Struthio camelus) male standing in landscape, Table Mountain National Park, Western Cape, South Africa

Ann & Steve Toon / Atlantic salmon (Salmo salar) leaping on upstream migration, River Tyne, Hexham, Northumberland, UK, November

Ann & Steve Toon / Spring wildflowers, Papkuilsfontein farm, Nieuwoudtville, Northern Cape, South Africa, September 2015

Ann & Steve Toon / Spring wildflowers, Papkuilsfontein farm, Nieuwoudtville, Northern Cape, South Africa, September 2015

Ann & Steve Toon / Spring wildflowers, Papkuilsfontein farm, Nieuwoudtville, Northern Cape, South Africa, September 2015

Ann & Steve Toon / White rhinoceros calf (Ceratotherium simum), Mkhuze Game Reserve, KwaZulu-Natal, South Africa, May