Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

Anup Shah / African elephant (Loxodonta africana) near water approaching - remote camera. Masai Mara National Reserve, Kenya.

Anup Shah / Olive baboon (Papio anubis) male approaching with curiosity - remote camera. Masai Mara National Reserve, Kenya.

Anup Shah / African elephant (Loxodonta africana) close up of leag approaching - remote camera. Masai Mara National Reserve, Kenya. August 2015.

Anup Shah / Mixed herd of Common or Plains zebra (Equus quagga burchellii) and Eastern white-bearded wildebeest (Connochaetes taurinus) - remote camera. Masai Mara National Reserve, Kenya..

Anup Shah / Vervet monkey (Cercopithecus aethiops) male scratching itself - remote camera . Masai Mara National Reserve, Kenya..

Anup Shah / African elephant (Loxodonta africana) approaching camera with curiosity, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Lion (Panthera leo) male approaching, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) feeding on loose soil for its minerals, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) family herd feeding on loose soil for its minerals, with dramatic stormy skies behind, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / African elephants (Loxodonta africana) family herd feeding on loose soil for its minerals, with dramatic stormy skies behind, Maasai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Mountain gorilla (Gorilla gorilla beringei) infant portrait, member of 'Humba' group. Virunga National Park, Democratic Republic of Congo, March.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 41 years grooms her daughter 'Golden' aged 14 years while their infants 'Gizmo' aged 3 years and 'Glamma' aged 9 months sit b...

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juvenile approaching with curiosity. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langurs (Semnopithecus dussumieri) rear view sitting, gathered on sandstone cliffs . Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langurs (Semnopithecus dussumieri) walking along a ledge . Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langurs (Semnopithecus dussumieri) sitting on sandstone cliffs . Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) group resting on rocky ledge. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langurs (Semnopithecus dussumieri) group of females feeding on potatoes left as an offering at a temple. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) female grooming a domestic dog. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juvenile approaching with curiosity. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langurs (Semnopithecus dussumieri) playing on the cliff tops at sunrise . Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juvenile playing silhouetted at sunrise, Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juveniles playing. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) females hugging to reinforce rank. Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juveniles playing at sunrise, Jodhpur, Rajasthan, India. March.

Anup Shah / Southern plains grey langur / Hanuman langur (Semnopithecus dussumieri) juveniles playing . Jodhpur, Rajasthan, India. March.

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River. Masai Mara National Reserve, Kenya.

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River. Masai Mara National Reserve, Kenya.

Anup Shah / Elephant (Loxodonta africana) calves playing together, Maasai Mara National Reserve, Kenya. December

Anup Shah / Elephant (Loxodonta africana) calves playing together . Maasai Mara National Reserve, Kenya. December

Anup Shah / Lioness with a playful cubs aged 3-6 months (Panthera leo). Maasai Mara National Reserve, Kenya. December

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) young baby of adolescent female 'Golden' suckling. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) male 'Faustino' aged 22 years walking along a track preparing to display. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) adult male 'Frodo' aged 35 years portrait. Gombe National Park, Tanzania.

Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding her daughter 'Glitter's' baby daught aged 2 months and her own infant son aged 2 years. With 'Glitter' aged...

Anup Shah / African elephants (Loxodonta africana) wallowing at a waterhole, wide angle view. Maasai Mara National Reserve, Kenya.

Anup Shah / Marabou storks (Leptoptilos crumeniferus) feeding on catfish in waterhole, wide angle view. Maasai Mara National Reserve, Kenya.

Anup Shah / African elephant (Loxodonta africana) walking toward camera, Maasai Mara National Reserve, Kenya.

Anup Shah / Cheetah (Acinonyx jubatus) male juvenile aged about 18 months approaching a kill, wide angle view. Maasai Mara National Reserve, Kenya.

Anup Shah / African elephants (Loxodonta africana) gathering at a waterhole, wide angle view. Maasai Mara National Reserve, Kenya.

Anup Shah / Dusky leaf monkey (Trachypithecus obscurus) juvenile yawning . Khao Sam Roi Yot National Park, Thailand. March 2015.

Anup Shah / Dusky leaf monkeys (Trachypithecus obscurus) in tree, one holding baby . Khao Sam Roi Yot National Park, Thailand.

Anup Shah / Dusky leaf monkey (Trachypithecus obscurus) baby playing . Khao Sam Roi Yot National Park, Thailand.

Anup Shah / Long-tailed macaque (Macaca fascicularis) juvenile sticking tongue out. Khao Sam Roi Yot National Park, Thailand.

Anup Shah / Tana mangabey (Cercocebus galeritus) mother grooms her baby of between -3 months old watched by a juvenile. Tana River Forest, South eastern Kenya.

Anup Shah / Lioness (Panthera leo) rejecting a male's advances Maasai Mara National Reserve, Kenya.

Anup Shah / Lion (Panthera leo) juveniles playing. Maasai Mara National Reserve, Kenya.