Robert Thompson / A fallen Sweet chestnut fruit (Castanea sativa) lying on the woodland floor, with a tree trunk covered in Textured lungwort (Lobaria scrobiculata) lichen, UK. October.
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Brian Bevan / Hedgehog (Erinaceus europaeus) walking through Autumn leaves, Bedfordshire, UK, November.
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Markus Varesvuo / Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October.
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Guy Edwardes / Ancient Roman bridge and autumn leaves, Allt da Ghob Waterfall, Glen Lyon, Perthshire, Scotland, UK. October.
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Guy Edwardes / Clustered bonnet fungi (Mycena inclinata) in broadleaved woodland, New Forest National Park, Hampshire, England, UK. Focus stacked image. November.
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Neil Aldridge / Tracking shot over a river, Dartmoor, Devon, England, UK, November.
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Simon Littlejohn / Adult female European otter (Lutra lutra) standing with its pup (approximately 14 weeks old) on the edge of the river, the pup raises its front paws onto mum's neck and the female then heads into the...
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Simon Littlejohn / Adult female European otter (Lutra lutra) guarding and grooming its pup on riverbank as it eats a fish, River Stour, Dorset, UK, September.
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Brian Bevan / Harvest mouse (Micromys minutus) climbing down blackberries on branch, Bedfordshire, UK, September. Controlled conditions.
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Brian Bevan / Melanistic grey squirrel (Sciurus carolinensis) searching for, finding and eating chestnuts amongst autumn leaves, Bedfordshire, UK, October.
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Pal Hermansen / An aerial shot of an autumnal birch forest swaying in strong wind, Rondane, Viken, Norway, September.
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Guy Edwardes / Spectacular rustgill (Gymnopilus junonius) at night with rain, New Forest National Park, Hampshire, England, UK. October.
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Guy Edwardes / Magpie inkcap fungus (Coprinopsis picacea). New Forest National Park, England, UK. October.
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Ben Hall / Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October.
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Tony Heald / Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October.
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Edwin Giesbers / Three Long-eared owls (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Birds in their Habitat Category winner, Expos Photos Aves Emotion'Ailes Competition 2020.
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Edwin Giesbers / Long-eared owl (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Second Prize Winner, Birds in their Environment Category, Bird Photographer of the Year (BPOTY) 2019...
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Marie Read / Blue jay (Cyanocitta cristata) in flight carrying an acorn that it will cache for winter food supply, New York, USA, October.
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Danny Green / Wolf (Canis lupus) standing at woodland edge. Finland, April.
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Guy Edwardes / Autumn colour around the Falls of Clyde, Lanarkshire, Scotland, UK, October 2006.
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Guy Edwardes / Tarn Hows, Lake District National Park, Cumbria, England, November 2007
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Jack Dykinga / Arizona sycamore (Platanus wrightii) roots and leaves amongst pebbles. In riparian area, Chiricahua Mountains, Coronado National Forest, Arizona, USA. November.
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Edwin Giesbers / Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October.
Guy Edwardes / Corfe Castle in early morning mist, Dorset, England, UK, October 2018.
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Andres M. Dominguez / Crested tit (Lophophanes cristatus) Sierra de Grazalema Natural Park. Southern Spain. September.
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Andres M. Dominguez / Cork trees (Quercus suber) stripped of bark, in forest. Los Alcornocales Natural Park, Cortes de la Frontera, southern Spain, November.
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Andres M. Dominguez / Hawfinch (Coccothraustes coccothraustes) perched on branch amongst Hawthorn (Crataegus sp) berries. Sierra de Grazalema Natural Park, Cadiz, Spain. February.
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Ingo Arndt / Aerial shot over a forest of Southern beech trees (Nothofagus) in autumn, Torres del Paine National Park, Patagonia, Chile, April, 2018.
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Jack Dykinga / Rainbow over Long Canyon with fall coloured Cottonwood (Populus) trees against canyon walls, Burr Trail, Grand Staircase-Escalante National Monument, Utah, USA, October 2018.
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Jack Dykinga / Waterfall over sandstone with cottonwood tree, Long Canyon , Burr Trail, Grand Staircase-Escalante National Monument, Utah, USA. October.
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Jack Dykinga / Morning light with autumn coloured Cottonwoods (Populus sp) trees lining the river. Valley of the Gods, Colorado Plateau, of the Great Basin Desert, Utah, USA, November 2018.
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John Shaw / Hybrid Poplar tree plantation in autumn (Populs sp.). Boardman Tree Farm, near Irrigon, Oregon, USA. October.
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Nick Hawkins / Atlantic salmon (Salmon salar) under river ice. Quebec, Canada. October.
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Marie Read / Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October.
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Marie Read / Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October.
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Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September
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Bruno D'Amicis / Red fox (Vulpes vulpes) walking along fallen trunk of old Beech (Fagus sylvatica) tree, Coppo del Principe old-growth beech forest in autumn. Abruzzo, Lazio and Molise National Park / Parco Nazionale...
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Sven Zacek / Bog surrounding river in morning light. Emajoe Suursoo Nature Reserve, Tartumaa, Estonia. October 2015.
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Sven Zacek / Flooded meanders of Halliste River, aerial view. Soomaa National Park, Parnumaa, Western Estonia. October 2017.
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Philippe Clement / Two Lynx (Lynx lynx) kittens play fighting in front of den entrance, Bavarian Forest National Park, Germany, October. Captive.
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TJ Rich / Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
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David Allemand / Birch tree (Betula sp) next to waterfall, Iceland. Runner-Up in the Plants category of the International OasisPhotoContest 2018
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Erlend Haarberg / Red squirrel (Sciurus vulgaris) hiding in a tree trunk, Klaebu, Norway, September.
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Guy Edwardes / Waterfall in Fjaorargljufur canyon, Kirkjubaejarklaustur, Iceland, August 2017.
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Guy Edwardes / View of hikers from behind the Pericnik Waterfall, Slovenia, October 2018.
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Bernard Castelein / Beech tree (Fagus sylvatica) showing roots, Peerdsbos, Brasschaat, Belgium