Charlie Hamilton James / Kingfisher (Alcedo atthias) adult male diving into water, Halcyon River, Gloucestershire, England.
Vishal Jadhav / Great hornbill (Buceros bicornis) pair performing courtship feeding ritual, passing berry from beak to beak, then female eats berry, Maharashtra, India, February.
Oliver Wright / A female kingfisher (Alcedo atthis) fishing through an ice hole in winter, emerging with prey in beak. Leeds, Yorkshire, UK. January.
Markus Varesvuo / Robin (Erithacus rubecula) carrying a red berry in its beak as it takes off, Parainen Uto, Finland. October.
Simon Littlejohn / Common kingfisher (Alcedo atthis) perched outside its nest, opening its wings and beak in an aggressive display to a Common blackbird (Turdus merula) which eventually flies away, River Stour, Dorset,...
Doug Gimesy / Portrait of Emu (Dromaius novaehollandiae).
Kangaroo Island, South Australia, Australia.
Pal Hermansen / Close up of a Emperor penguin (Aptenodytes forsteri) face looking around and towards camera, Snow Hill Island, Antarctica, October.
Edwin Giesbers / Dalmatian pelican (Pelicanus crispus) group squabbling over fish, Lake Kerkini, Greece.
Guy Edwardes / Long-billed curlew (Numenius americanus) taking off along shoreline. Fort De Soto Park, Florida, USA. April.
Guy Edwardes / Dalmatian pelican (Pelecanus crispus), Lake Kerkini, Greece
Markus Varesvuo / Red-throated Diver (Gavia stellata) portrait, Iceland, June.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick looking upwards to parents beak. Atka Bay, Antarctica. August. Winner of the Portfolio category of the Wildlife Photographer of the Year Awards 2019.
Jack Dykinga / Greater roadrunner (Geococcyx californianus) running to feed brood with Lizard prey in beak. Catalina State Park, Sonoran Desert, Arizona, USA. June.
George Sanker / Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2.
Oscar Dewhurst / Great bittern (Botaurus stellaris) with recently caught fish prey in beak. Suffolk, UK. November.
SCOTLAND: The Big Picture / Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October..
Marie Read / American oystercatcher (Haematopus palliatus) adult feeding chick, Long Island, New York, USA. June.
Marie Read / American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April.
Marie Read / California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Marie Read / Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October.
Marie Read / Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions.
Marie Read / Great blue heron (Ardea herodias) in breeding plumage, head and shoulders showing plumes on crown and neck, Montezuma National Wildlife Refuge, New York, USA, May.
Marie Read / Ruby-throated Hummingbird (Archilocus colubris), male flying in to feed from cardinal flowers (Lobelia cardinalis) New York, USA. August.
John Chan / Male Anna's hummingbird (Calypte anna) cleaning its beak on a twig, Southern California, USA, January.
George Sanker / Greater yellowlegs (Tringa melanoleuca) standing on rock. Acadia National Park, Maine, USA. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August. Sequence 1/2.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick begging for food, on parent's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) egg hatching on father's feet. Atka Bay, Antarctica. August.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri) chick aged six to eight weeks stretching whilst on brooding parent's feet. Adult looking down at chick. Atka Bay, Antarctica. September.
Stefan Christmann / Emperor penguin (Aptenodytes forsteri), two chicks aged six to eight weeks brooding on parents' feet, one vocalising. Atka Bay, Antarctica. September.
Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.
Paul Hobson / Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May.
Paul Hobson / Spoonbill (Platalea leucorodia) preening, close up. Hungary. May.
Kerstin Hinze / Red crossbill (Loxia curvirostra) feeding from a spruce cone, Bavarian Forest National Park, Germany, October.
Sven Zacek / Ural owl (Strix uralensis) female on nest, male landing with twig in beak. Tartumaa, Southern Estonia. April.
Sven Zacek / Black woodpecker (Dryocopus martius) excavating nest hole. Valgamaa, Southern Estonia. April.
Sven Zacek / Ural owl (Strix uralensis) female flying with prey in beak. Flying back to nest in evening light. Tartumaa, Southern Estonia. May.
Sven Zacek / Great bittern (Botaurus stellaris) with fish in beak, caught in stream. Laanemaa, Western Estonia. March.
Tui De Roy / Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos.
Tui De Roy / Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos.
Konrad Wothe / Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador.
Steven David Miller / Emu (Dromaius novaehollandiae) juvenile with beak open to catch a leaf, Cleland Wildlife Park, South Australia, March. Captive.
Guy Edwardes / Little bee-eater (Merops pusillus) vocalising whilst perched on branch, Khwai, Botswana.
Claudio Contreras / Caribbean Flamingo (Phoenicopterus ruber) sleeping in the breeding colony, brooding egg, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June, Finalist in the Portfolio Category of the Terr...