Chien Lee / Minden / Black-headed Cat Snake (Boiga nigriceps) swallowing alive Harlequin Flying Tree Frog (Rhacophorus pardalis) prey, Kubah National Park, Sarawak, Malaysia, sequence 1 of 4
Sergey Gorshkov / Minden / Northern Goshawk (Accipiter gentilis) feeding on Black Grouse (Tetrao tetrix) female in winter, Finland
Winfried Wisniewski / European Bee-eater (Merops apiaster) pair, Pleven, Bulgaria
Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) defacating by hanging from thumbs the 'right way up', Yarra Bend Park, Victoria, Australia, April.
Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) aggressive mating behaviour (wrestling, biting) male eventually moves away, Yarra Bend Park, Victoria, Australia, April.
Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) male involved in courtship / mating behaviour including licking female's genitalia; male grooming his own penis, bickering and female climbing away, Yar...
Neil Challis / Little grebe (Tachybaptus ruficollis) chasing a small fish jumping out of the water, before finally catching then eating it, Western Thailand.
Neil Challis / Two juvenile Long-tailed macaques (Macaca fascicularis) play fighting in a tree during a shower, Western Thailand.
Neil Challis / Weaver ants (Oecophylla sp.) attacking and pulling the legs of a Formicine ant (Polyrhachis proxima) on the forest floor, Western Thailand.
Neil Challis / Weaver ants (Oecophylla sp.) pulling apart a Kapok bug (Probergrothius nigricornis) and carrying it back to their nest, Western Thailand.
Marie Read / Common loon (Gavia immer) adult swimming with a chick resting on its back, Presque Isle County, Michigan, USA, June.
Marie Read / American bittern (Botaurus lentiginosus) calling / booming in a cattail marsh to mark territory, Salt Alkaline Lake, Kidder County, North Dakota, USA, June.
Lukas Schaefer / Rock ptarmigan (Lagopus muta) close up preening, Val Venosta, Italy, October.
Vishal Jadhav / Exhausted Short-eared owl (Asio flammeus) surrounded by House crow (Corvus splendens) group at dawn, one crow dives down to attack, Maharashtra, India, April.
Vishal Jadhav / Short-eared owl (Asio flammeus) stood surrounded by a group of House crows (Corvus splendens) trying to find weakness by snapping at its feathers, Maharashtra, India, April.
Vishal Jadhav / Red-breasted flycatcher (Ficedula parva) male hunting flying insects from a rock, Maharashtra, India, February.
Vishal Jadhav / Great hornbill (Buceros bicornis) pair performing courtship feeding ritual, passing berry from beak to beak, then female eats berry, Maharashtra, India, February.
Nick Upton / Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK,...
Vishal Jadhav / Asian green bee-eater (Merops orientalis) hunting insects from perch in the grass, Maharashtra, India, July.
Vishal Jadhav / Indian jackal (Canis aureus indicus) pair howling, Maharashtra, India, September.
Vishal Jadhav / Indian jackal (Canis aureus indicus) female grooming male's head, Maharashtra, India, September.
Noemie Stroh / Green turtles (Chelonia mydas) mating on coral reef, Rocky Island, Red Sea, Egypt, June.
Matt Roseveare / Caddisfly larva (Trichoptera) building armoured casing / case with pebbles on riverbed, Cornwall, UK, April. Controlled conditions.
Matt Roseveare / White-throated dipper (Cinclus cinclus) de-shelling caddisfly larva (Trichoptera) then eating it, River Kennal, Cornwall, UK, February.
Matt Roseveare / Parson's chameleon (Calumma parsonii) catching insect with tongue, Eastern rainforests of Madagascar, September.
Oliver Wright / A female kingfisher (Alcedo atthis) fishing through an ice hole in winter, emerging with prey in beak. Leeds, Yorkshire, UK. January.
Oliver Wright / A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January.
Oscar Dewhurst / Mandarin duck (Aix galericulata) drake swimming on pond, backlit by rising sun and bokeh effect, Richmond Park, London, UK. February.
David Pattyn / Pair of Great crested grebes (Podiceps cristatus) performing courtship dance, The Netherlands, Europe. March.
David Pattyn / Coots (Fulica Atra) fighting in early morning light. Valkenhorst Nature Reserve, The Netherlands, Europe. April.
David Pattyn / Pair of Great crested grebes (Podiceps cristatus) courtship dance display, The Netherlands, Europe. March.
Brian Bevan / Yellowhammer (Emberiza citronella) male singing from bramble then flies away, Bedfordshire, UK, June.
Brian Bevan / Common Kingfisher (Alcedo atthis) male dives then catches loach fish, Bedfordshire, UK, April.
Brian Bevan / Common Kingfisher (Alcedo atthis) male fishing from blackthorn branch, catches then eats fish, Bedfordshire, UK, April.
Brian Bevan / Great crested grebe (Podiceps cristatus) courtship weed dance, Bedfordshire, UK, April.
Eric Baccega / Male Chimpanzee (Pan troglodytes troglodytes) trying to catch fallen fruits using stick as a tool, Conkouati-Douli National Park, Republic of Congo, Africa.
Lewis Jefferies / Common octopus (Octopus vulgaris) changing colour as a threat display, Maldives, Indian Ocean.
Lewis Jefferies / Blackfinned anemonefish (Amphiprion nigripes) family seeks shelter in safety of Anemone's stinging tentacles, South Ari Atoll, Maldives, Indian Ocean.
Tui De Roy / Marine iguana (Amblyrhynchus cristatus) basking on driftwood, Fernandina Island, Galapagos, South America.
Henley Spiers / Striped marlin (Kajikia audax) striking a shoal of baitfish during the largest known seasonal aggregation of Marlin, Magdalena Bay, Baja California Sur, Mexico, Pacific Ocean. November 2019
Henley Spiers / Striped marlin (Kajikia audax) and California sea lion (Zalophus californianus) competing for food as they hunt down baitfish, Magdalena Bay, Baja California Sur, Mexico, Pacific Ocean.