Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) defacating by hanging from thumbs the 'right way up', Yarra Bend Park, Victoria, Australia, April.
Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) aggressive mating behaviour (wrestling, biting) male eventually moves away, Yarra Bend Park, Victoria, Australia, April.
Doug Gimesy / Grey-headed flying-fox (Pteropus poliocephalus) male involved in courtship / mating behaviour including licking female's genitalia; male grooming his own penis, bickering and female climbing away, Yar...
Marie Read / Common loon (Gavia immer) adult swimming with a chick resting on its back, Presque Isle County, Michigan, USA, June.
Marie Read / American bittern (Botaurus lentiginosus) calling / booming in a cattail marsh to mark territory, Salt Alkaline Lake, Kidder County, North Dakota, USA, June.
Lukas Schaefer / Rock ptarmigan (Lagopus muta) close up preening, Val Venosta, Italy, October.
Vishal Jadhav / Exhausted Short-eared owl (Asio flammeus) surrounded by House crow (Corvus splendens) group at dawn, one crow dives down to attack, Maharashtra, India, April.
Vishal Jadhav / Short-eared owl (Asio flammeus) stood surrounded by a group of House crows (Corvus splendens) trying to find weakness by snapping at its feathers, Maharashtra, India, April.
Vishal Jadhav / Red-breasted flycatcher (Ficedula parva) male hunting flying insects from a rock, Maharashtra, India, February.
Vishal Jadhav / Great hornbill (Buceros bicornis) pair performing courtship feeding ritual, passing berry from beak to beak, then female eats berry, Maharashtra, India, February.
Nick Upton / Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK,...
Vishal Jadhav / Asian green bee-eater (Merops orientalis) hunting insects from perch in the grass, Maharashtra, India, July.
Vishal Jadhav / Indian jackal (Canis aureus indicus) pair howling, Maharashtra, India, September.
Vishal Jadhav / Indian jackal (Canis aureus indicus) female grooming male's head, Maharashtra, India, September.
Noemie Stroh / Green turtles (Chelonia mydas) mating on coral reef, Rocky Island, Red Sea, Egypt, June.
Matt Roseveare / Caddisfly larva (Trichoptera) building armoured casing / case with pebbles on riverbed, Cornwall, UK, April. Controlled conditions.
Matt Roseveare / White-throated dipper (Cinclus cinclus) de-shelling caddisfly larva (Trichoptera) then eating it, River Kennal, Cornwall, UK, February.
Matt Roseveare / Parson's chameleon (Calumma parsonii) catching insect with tongue, Eastern rainforests of Madagascar, September.
Oliver Wright / A female kingfisher (Alcedo atthis) fishing through an ice hole in winter, emerging with prey in beak. Leeds, Yorkshire, UK. January.
Oliver Wright / A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January.
Oscar Dewhurst / Mandarin duck (Aix galericulata) drake swimming on pond, backlit by rising sun and bokeh effect, Richmond Park, London, UK. February.
David Pattyn / Pair of Great crested grebes (Podiceps cristatus) performing courtship dance, The Netherlands, Europe. March.
David Pattyn / Coots (Fulica Atra) fighting in early morning light. Valkenhorst Nature Reserve, The Netherlands, Europe. April.
David Pattyn / Pair of Great crested grebes (Podiceps cristatus) courtship dance display, The Netherlands, Europe. March.
Brian Bevan / Yellowhammer (Emberiza citronella) male singing from bramble then flies away, Bedfordshire, UK, June.
Brian Bevan / Common Kingfisher (Alcedo atthis) male dives then catches loach fish, Bedfordshire, UK, April.
Brian Bevan / Common Kingfisher (Alcedo atthis) male fishing from blackthorn branch, catches then eats fish, Bedfordshire, UK, April.
Brian Bevan / Great crested grebe (Podiceps cristatus) courtship weed dance, Bedfordshire, UK, April.
Eric Baccega / Male Chimpanzee (Pan troglodytes troglodytes) trying to catch fallen fruits using stick as a tool, Conkouati-Douli National Park, Republic of Congo, Africa.
Lewis Jefferies / Common octopus (Octopus vulgaris) changing colour as a threat display, Maldives, Indian Ocean.
Lewis Jefferies / Blackfinned anemonefish (Amphiprion nigripes) family seeks shelter in safety of Anemone's stinging tentacles, South Ari Atoll, Maldives, Indian Ocean.
Tui De Roy / Marine iguana (Amblyrhynchus cristatus) basking on driftwood, Fernandina Island, Galapagos, South America.
Henley Spiers / Striped marlin (Kajikia audax) striking a shoal of baitfish during the largest known seasonal aggregation of Marlin, Magdalena Bay, Baja California Sur, Mexico, Pacific Ocean. November 2019
Henley Spiers / Striped marlin (Kajikia audax) and California sea lion (Zalophus californianus) competing for food as they hunt down baitfish, Magdalena Bay, Baja California Sur, Mexico, Pacific Ocean.
Terry Whittaker / Two Arctic fox (Vulpes lagopus) cubs playing together near den, Hornvik, Hornstrandir Nature reserve, Iceland, July.
Terry Whittaker / Arctic fox (Vulpes lagopus) cubs playing with feather, playfighting and chasing each other near den, Hornvik, Hornstrandir Nature reserve, Iceland, July.
Simon Littlejohn / Female American mink (Neogale vison) entering a Kingfisher (Alcedo atthias) nest and taking a chick, Stour River, Dorset, UK, July.
Simon Littlejohn / Common kingfisher (Alcedo atthis) perched outside its nest, opening its wings and beak in an aggressive display to a Common blackbird (Turdus merula) which eventually flies away, River Stour, Dorset,...
Klein & Hubert / Nile crocodile (Crocodylus niloticus) close up of jaw and teeth, Chobe River, Botswana, Africa.
Klein & Hubert / Cape buffalo (Syncerus caffer) with Yellow-billed oxpckers (Buphagus africanus) drinking at waterhole in dry season, South Luangwa National Park, Zambia, Africa.
Klein & Hubert / African elephant (Loxodonta africana) calf playing in savanna, Kenya, Africa.
Terry Whittaker / White-throated dipper (Cinclus cinclus) singing on old bicycle wheel, River Mersey, Greater Manchester, UK.
Terry Whittaker / White-throated dipper (Cinclus cinclus) returning to newly-fledged chick to feed it before flying away, River Mersey, Greater Manchester, UK.
Kim Taylor / Song thrush (Turdus philomelos) cracking a Garden snail (Cornu aspersum) on stone 'anvil', Surrey, UK, June.
Kim Taylor / Female Red mason bee (Osmia bicornis) stocking her nest in a glass tube with nectar and pollen before laying an egg, followed by her sealing the cell with mud, Surrey, UK, April.