Vishal Jadhav / European roller (Coracias garrulus) perched on a rock preening its feathers, Maharashtra, India, February.
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Marie Read / American bittern (Botaurus lentiginosus) calling / booming in a cattail marsh to mark territory, Salt Alkaline Lake, Kidder County, North Dakota, USA, June.
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Vishal Jadhav / Indian jackal (Canis aureus indicus) pair howling, Maharashtra, India, September.
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Claudio Contreras / Laysan albatross (Phoebastria immutabilis) calling mate,Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
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Ismaele Tortella / Eurasian otter (Lutra lutra) swimming with snow on its back, calling, Norway
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Alex Mustard / Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home, a Haddon's sea anemone (Stichoda...
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Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male from branch, flies off, Bedfordshire, England, UK, May.
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Brian Bevan / Male European cuckoo (Cuculus canorus) calling to another male, before flying off in response to a female, Bedfordshire, England, UK, May.
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Danny Green / Red deer stag (Cervus elaphus) roaring. Bradgate Park, Leicestershire, UK, October.
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Erlend Haarberg / Black grouse (Lyrurus tetrix), male calling early in the morning, Norway. May.
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Markus Varesvuo / Red-throated Diver (Gavia stellata) calling, Iceland, June.
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Marie Read / Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion.
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Marie Read / American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May.
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Marie Read / Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
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Marie Read / Red-winged Blackbird (Agelaius phoeniceus) male displaying in cattail marsh, Viera Wetlands, Brevard County, Florida, USA, January.
Klein & Hubert / Grey wolf (Canis lupus) female and two cubs, one howling. Controlled conditions
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Klein & Hubert / Wolf (Canis lupus) puppy, age one month, Rocky Mountains, USA. Controlled conditions.
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Stefan Christmann / Adelie penguin (Pygoscelis adeliae) calling during courtship, Atka Bay, Queen Maud Land, Antarctica, December.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri) male feeding chick aged a few days. Atka Bay, Antarctica. August. Sequence 1/2.
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Stefan Christmann / Emperor penguin (Aptenodytes forsteri), group of three males incubating, one with begging chick. Atka Bay, Antarctica. August.
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Guy Edwardes / Marsh Frog (Pelophylax ridibundus) vocalising with bokeh effect in the background, Danube Wetlands, Bulgaria
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Guy Edwardes / Marsh Frog (Pelophylax ridibundus) vocalising with bokeh effect in the background, Danube Wetlands, Bulgaria
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David Perpinan / Male Mediterranean tree frog (Hyla meridionalis) calling, showing vocal sac inflating, Barcelona, Spain, May. Filmed at night with an infrared camera.
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Philippe Clement / Territorial Common raven (Corvus corax) on cliff top, tearing out vegetation, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Philippe Clement / Common raven (Corvus corax) calling from top of sea cliff, Eshaness, Northmavine, Shetland Islands, Scotland, UK, May.
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Konrad Wothe / Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June.
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
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Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.
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David Perpinan / Oystercatcher (Haematopus ostralegus) incubating eggs on nest, alarm calling as a predator flies overhead, Scotland,UK, June.
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Tui De Roy / Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
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Fred Olivier / Emperor penguin (Aptenodytes forsteri) chick begging for food from parent, Adelie Land, Antarctica, January.
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Fred Olivier / Emperor penguin (Aptenodytes forsteri) chicks calling, with another small chick arriving and falling over, Adelie Land, Antarctica, January.
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Yashpal Rathore / Jayarami Bush frog (Raorchestes jayarami), inflating vocal sac, calling to attract mate, Endemic to Western Ghats. Anamalai Wildlife Sanctuary, India.
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Markus Varesvuo / Japanese / Red-crowned crane (Grus japonicus) two calling and dancing, part of bonding and courtship display, Hokkaido Japan February
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Eric Baccega / Pack of Grey wolves (Canis lupus) howling in the snow, captive. Bavarian Forest National Park, Germany, March. Captive.
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Ernie Janes / RF- Lesser black-backed gulls (Larus fuscus) calling from shoreline, beaks open, England, UK, May. (This image may be licensed either as rights managed or royalty free.)
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Neil Aldridge / Lion cub (Panthera leo) calling to its mother from the raised deck of a tourist lodge in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable species.
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Mark Hamblin / Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016.
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Alex Hyde / Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
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Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 20 years, females Nuru' aged 22 years and 'Nasa' aged 24 years calling. Gombe National Park, Tanzania. June 2012.
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Klein & Hubert / Glaucous gull (Larus hyperboreus) standing on edge of ice calling, Svalbard, Norway
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Bence Mate / Montezuma oropendola (Psarocolius montezuma) two interacting, Costa Rica.
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Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years calling while holding her baby granddaughter aged 2 months and her own infant son aged 2 years. Gombe National Park,...
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Mark Bowler / Black spider monkey (Ateles chamek) calling, captive, Peru.
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Cyril Ruoso / Atlantic puffin (Larus tridactyla) seen from burrow, Iceland, July.
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Yukihiro Fukuda / Japanese macaque (Macaca fuscata fuscata) rare white furred baby calling, Jigokudani Valley, Nagano Prefecture, Japan. June.
