Ross Hoddinott / Red fox (Vulpes vulpes) vixen yawning in the morning light (captive). Lifton, Devon, UK, May

Anup Shah / African Lion (Panthera leo) juvenile reaching out with curiosity, Masai Mara National Reserve, Kenya. March

Edwin Giesbers / Saltwater crocodile (Crocodylus porosus) close up portrait, Sarawak, Borneo, Malaysia

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tata' and her unnamed baby aged 2-3 months portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010. Reh...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) mature male 'Doyok's' hand holding a liana(Pongo pygmaeus wurmbii). Pondok Tanggui, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Yuni' and her baby aged 3-6 months - portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. July 2010. Rehabilit...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Tutut' sitting with her sleeping baby 'Thor' aged 8-9 months. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. July...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) sub-adult male 'Oman' sitting on a clump of vegetation in the river feeding - wide angle perspective. Camp Bulu, Lamandau Nature Reserve, Central Kalimantan,...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) young mature male 'Gary' aged 16 years portrait. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. July 2010. Rehabilitated a...

Anup Shah / Bornean Orangutan (Pongo pygmaeus wurmbii) female 'Peta' carrying her baby daughter 'Petra' aged 12 months. Camp Leakey, Tanjung Puting National Park, Central Kalimantan, Borneo, Indonesia. June 2010....

Anup Shah / African lion (Panthera leo) cubs aged 6-9 months approaching with curiosity watched by their mother, Masai Mara National Reserve, Kenya. February

Anup Shah / African elephant (Loxodonta africana) group leaving a waterhole - wide angle perspective. Masai Mara National Reserve, Kenya. March

Anup Shah / African elephant (Loxodonta africana) group drinking from waterhole - wide angle perspective. Masai Mara National Reserve, Kenya. March

Anup Shah / White-backed vultures (Gyps africanus) and Black-back jackal (Canis mesomelas) feeding from the carcass of a Cape buffalo (Cyncerus caffer) - wide angle perspective. Masai Mara National Reserve, Kenya...

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective. Masai Mara National Reserve, Kenya. August

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective, Masai Mara National Reserve, Kenya. August

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective, Masai Mara National Reserve, Kenya. October

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective, Masai Mara National Reserve, Kenya. October

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective Masai Mara National Reserve, Kenya. October

Anup Shah / Eastern White-bearded Wildebeest herd (Connochaetes taurinus) and Common / Plains zebra (Equus quagga burchellii) running away from the Mara River after crossing - wide angle perspective. Masai Mara N...

Anup Shah / Common / Plains Zebra (Equus quagga burchellii) herd running away from the Mara River - wide angle perspective, Masai Mara National Reserve, Kenya. September 2009.

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River - wide angle perspective, Masai Mara National Reserve, Kenya. September 2009.

Anup Shah / Thomson's gazelles (Eudorcas thomsonii) following a herd of Wildebeest (Connochaetes taurinus) as they cross the Mara River, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Cape / African Buffalo (Syncerus caffer) approaching with curiosity, herd in the background, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Cape / African Buffalo (Syncerus caffer) approaching camera with curiosity, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running, rear view, wide angle perspective, Masai Mara National Reserve, Kenya. September

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd running - wide angle perspective, Masai Mara National Reserve, Kenya. August 2009.

Anup Shah / Common / Plains Zebra (Equus quagga burchellii) close up view of under belly - wide angle perspective. Masai Mara National Reserve, Kenya. August

Anup Shah / Eastern White-bearded Wildebeest (Connochaetes taurinus) herd crossing the Mara River, close up of legs - wide angle perspective. Masai Mara National Reserve, Kenya. September 2009.

Anup Shah / Masai giraffe (Giraffa camelopardalis tippelskirchi) crossing the Mara River - wide angle perspective, Masai Mara National Reserve, Kenya. August 2009.

Nick Upton / Hoverfly (Myathropa florea), using proboscis to feed on pollen from anthers of Ivy flowers (Hedera helix). Wiltshire garden, UK, September

Fabio Liverani / Freshwater Crab (Potamon fluviatile) brandishing claws in defensive posture. Foreste Casentinesi National Park, Italy, July.

Fabio Liverani / Freshwater Crab (Potamon fluviatile) in a shallow stream - split perspective showing crab beneath and fluvial habitat. Foreste Casentinesi National Park, Italy, July.

Fabio Liverani / Freshwater Crab (Potamon fluviatile) in a shallow stream - split perspective of underwater and habitat. Foreste Casentinesi National Park, Italy, July.

Edwin Giesbers / Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January.

Edwin Giesbers / Wild Boar (Sus scrofa) males fighting in snow. The Netherlands, January.

Edwin Giesbers / Common Pheasant (Phasianus colchicus) on grassland. The Netherlands.

Edwin Giesbers / White-fronted Goose (Anser albifrons) in flight over grassland. The Netherlands.

Edwin Giesbers / Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January.

Edwin Giesbers / European Mouflon (Ovis musimon) male head profile. The Netherlands, January.

Edwin Giesbers / European Mouflon (Ovis musimon) male in snow. The Netherlands, January.

Edwin Giesbers / Whooper Swan (Cygnus cygnus) with its beak in its feathers. The Netherlands, January.

Edwin Giesbers / Grey Heron (Ardea cinerea) leaving footprints while walking through snow. The Netherlands, January.

Edwin Giesbers / Legs of a White Stork (Ciconia ciconia) on ice. The Netherlands, January.

Edwin Giesbers / Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June.

Edwin Giesbers / Whooper Swan (Cygnus cygnus) with its beak tucked under its wing. The Netherlands, June.

Edwin Giesbers / Mute Swan (Cygnus olor) standing on frosty grass. The Netherlands, March.

Edwin Giesbers / House Mouse (Mus musculus) peering through hole in cheese. Captive.